摘要
平贝母(Fritillaria usuriensis Maxim.)花药壁的发育为基本型。花粉母细胞减数分裂时的胞质分裂为连续型。四分孢子呈左右对称式排列。生殖细胞最初形成的壁为胼胝质的。散粉时的花粉粮为二细胞型。双珠被,薄珠心,倒生胚珠;胚囊发育属贝母型;成熟胚囊为七核组成的六细胞胚囊。双受精作用属于有丝分裂前配子融合类型。受精结束时未见到雌雄性核仁的融合。初生胚乳核的分裂先于合子的分裂;胚乳发育为核型;胚的发育属柳叶菜型。种子成熟时胚乳细胞未被吸收,没有胚的分化。
The development of the anther wall follows Basic--type. The cytokinesis at the time of pollen mother cell meiosis conforms to successive type. The arrangement of the microspores in the te- trad is referred to isobilateral. The primary wall between the generative cell and the vegetative cell is callose. The callose wall is easily detected under the fluorescence microscope. The mature pollen grain is 2--celled type. The ovule is bitegminous, tenui-nucellar and anatropous. The development of the female gametophyte follows Fritillaria-type. The mature embryo sac. consists of the six cells including the seven nuclei. The fertilization is referred to the premi- totic syngamy type. The fusion of the female and male nucleoli is not observed at the end of the fertilization. The division of the primary endosperm nucleus is earlier than that of the zygote. The development of the endosperm is referred to nuclear type. The division of the zygote is transverse of longitudinal, the development of the embryo conforms to Onagrad- type. When the seed is mature. the embryo is at the proembryo stage without differentiation and the endosperm cells are not absorbed.
基金
国家自然科学基金
关键词
平贝母
胚胎学
Fritillaria ussuriensis Maxim.
Anther
Pollen
Ovule
Embryo sac of Fritillaria-type
Fertilization
Embryo anl endosperm