摘要
革苞菊为雌雄异株。在雄花中 ,花药 4室 ,药壁发育为双子叶型 ,由表皮、药室内壁 ,一层中层和绒毡层组成。绒毡层于小孢子四分体时期开始变形 ,其细胞原生质体向药室中移动 ,为变形绒毡层。小孢子孢原为多细胞 ,小孢子母细胞减数分裂产生四面体型的小孢子四分体。四分体胞质分裂为同时型。成熟花粉 3-细胞型。单核期的小孢子出现壁发育不良和巨大及空花粉现象。在雌花中 ,胚珠是倒生的 ,单珠被 ,薄珠心 ,珠被于孢原期已发育完整。大孢子孢原单细胞。由孢原细胞直接发育形成大孢子母细胞。 4个大孢子直线型 ,蓼型胚囊。于成熟胚囊期观察到发育异常的胚囊。通过对胚囊发育过程中营养物质消长规律的研究 ,讨论了环境与发育的相关性问题。
Tugarinovia mongolica is dioecious plant.In male flowers,the anther has four chambers.The development of anther wall is dicotyledonous type which is composed of epidermis,endothecium,one layer of middle layer and amoeboid tapetum.The tapetum begins to vary at the stage of tetrad with the cytoplasm moving into the anther cell.The cytokinesis of microspore mother cell in meiosis is simuteneous,and the tetrad are tetrahedral.The mature pollen grain is 3 celled.The big,empty and abortive pollen grains are abserred at the stage of uninuclear microspore.In female flowers,Ovule is anatropous,unitegmic and tenuinuceuate with the development of integument finished at the stage of archespore.The megaspore mother cells are developed directly by a hypodermal archesporical cell.Which undergoes meitotic division and forms a linear tetrad.The embryo sac is of Polygonum type.Some abnormal mature embryo sacs are observed.The problem of the relations between the environment and the development is discussed on the foundation of the study of the histochemical changes.
出处
《西北植物学报》
CAS
CSCD
2000年第3期461-466,共6页
Acta Botanica Boreali-Occidentalia Sinica
基金
国家自然科学基金资助项目!( 3 9560 0 0 7)
关键词
革苞菊
孢子发生
配子体发育
环境
胚胎学
Tugarinovia mongolica
megasporogenesis and microsporogenesis
development of gametophytes
changes of histochemistry
