摘要
描述中生蝎蛉科Mesopanorpodidae Tillyard,1918化石2新属3新种:宽翅柱状蝎蛉Stylopanorpodes eurypterus gen.et sp.nov.,华美纺锤蝎蛉Netropanorpodes decorosus gen.et sp.nov.,多刺纺锤蝎蛉Netropanorpodes sentosus gen.et sp.nov.。所有化石标本均采于内蒙古宁城道虎沟中侏罗世九龙山组地层,模式标本现保存于首都师范大学生命科学学院。
Two new genera and three new species,Stylopanorpodes eurypterus gen.et sp.nov.,Netropanorpodes decorosus gen.et sp.nov.and Netropanorpodes sentosus gen.et sp.nov.,are described.All fossil specimens were collected from the Middle Jurassic Jiulongshan Formation of Inner Mongolia in China and deposited in the Capital Normal University.Stylopanorpodes gen. nov. Type species. Stylopanorpates euryptems gen. et sp Diagnosis. Moderate-sized insect, rostrum long and slender. Eyes large. Legs slender, covered with dense and tiny hairs, irregular mrrangement. The forewing broad, pterostigma not distinctly. Sc long, 2-branched, reaching the pterostigmal region. RI not branched, Rs 4-branched, forking before midwing, R4＋5 forking before R2＋3. M 4-branched, which forks later than Rs; M3＋4 forking before M1 ＋ 2. CuA and M fused basaUy for a long distance, cubito-merlizn Y-vein present. 2 crossveius between CuP and A1 present. Abdomen elongate, cylindrical, tapering apically, in the female 11 segments visible. Comparison. In the general venation scheme the Stylopanorpodes gen. nov. somewhat resembles Mesopanorpodes Tallyard, 1918 and Prochoristella Riek, 1953, but the new genus differs from the former by R4＋ forking before R2＋3, two cross-veins between 1A and CuP present, and differs from the latter by Rs forking before M, R2 ＋3 forking later, both R2 and R3 shorter, the cross-vein between CuA and CuP absent. The new genus also may be distinguished from other genera by the forking of Rs and M, and the cross-veins in apical area of Rs and M strongly degenerated. Etymology. The genus name is a combination of styl-（ Greek, meaning cylindrical ） and panorpodes （the holotype genus Mesopanorpodes in this family）.Stylopanorpodes eurypterus gen. et sp. nov. （Figs. 1- 4, 13） Etymology. Euryptems from Latin, means broad wing. Holotype. A well preserved female insect. Registration No： CaNU-M-NN-2006005. Locality and horizon. Daohugou Village, Ningcheng Gounty, Inner Mongolia, China; Jiulongshan Formation, Middle Jurassic. Description. Medium-sized insect, length l lmm. The specimen shows a lateral view of the whole insect. The left forewing spreading out ventrad to the body. The right forewing slightly overlapping the body. The hind wings folded together, venation not discernible. Vertex of the head raised, with extremely large protruding eyes at anterior of head. Rostrum slender and long. Antennae missing. Pronotum small, meso- and metanotum distinctly bigger. Legs slender and long, covered with dense and tiny hairs, irregularly arranged. Coxae very dear, which are divided distinctly into eucoxae and mera. Trochanters small. Hind tibia obviously longer than fernom, with 2 terminal calcars, about equal in length. Tarsus 5- segmented, basitarsus longer than others, about two times as long as the second tarsomere. Forewing length 11 mm, width 4 mm, apex evenly rounded, widest somewhat beyond the middle. Costal space expanded. Pterostigma not distinct. Sc long, 2- branched, extending to the pterostigmal region. R1 simple, not branched, reaching the wing margin, curving posteriorly near its temaination. Rs 4-branched, dividing into its two main branches before midwing, which forks ealier than M, R4＋ 5 forking before R2 ＋ 3- M arising from the base of the wing, 4-branched, M3＋4 forking before M1＋ 2, which forks after the forking of R4 ＋ 5. CuA and M fused basaUy for a long distance, cubito-medlan Y-vein present. CuP simple, well separated from CuA. 1A simple, 2 cross-veins between CuP and 1A present. 2A, 3A not preserved. The hind wing only partially preserved, venation not discernible. Abdomen long, cylindrical, with tapering termination, 11 segments visible. Basitergum fused to metathorax, slightly shortened, the 9th to 11 th segments abruptly more slender than 2-8, the cercus not dear. Netropanorpodes gen. nov. Type species. Netropanorpodes decorosus gen. et sp. nov. Diagnosis. Moderate-sized insect, rostrum long and slender, maxillary palp 4-segmented. Eyes large. Antennae filiform, many-segmented, with short, stout scape and long slender flagellum. Legs slender and long, covered with dense and tiny hairs, irregular arrangement. The forewing pterostigma dearly present. Sc long, 2-branched, reaching the pterostigmal region. R1 not branched, Rs 4-branched, forking nearly midwing, which forks nearly on the same level with M. M 4-branched, M3＋4 forking before M1 ＋2. CuA and M fused basaUy for a long distance, cubito-median Y-vein variable. 2 cross-veins between 1A and 2A present. The hind wing Sc shorter, not reaching the pterostigmal region. Abdomen fusiform, in the male 10 segments visible, and female 11 segments visible. Comparison. The new genus is siroilar to Stylopanorpodes gen. nov. However, the new genus forewing with dear pterostigma, Rs and M forked on the same level, two cross-veins between 1A and 2A present, the hind basitarsus longer, which is distinct from the later. The new genus also could be distinguished from other genera by Rs and M forked on the same level, mad the cross-veins in the area of Rs and M strongly degenerated. Etymology. The genus name is a combination of netro-（ Greek, meaning fusfform ） and panorpodes （ the holotype genus Mesopanorpodes in this family） .Netropanrorpodes Decorosus gen. et sp. nov. （Figs. 5- 8, 14） Etymology. Decorosus from Latin, means gorgeous. Holotype. A well preserved male insect. Registration No： CNU-M-NN-2006100. Locality and horizon. Daohugou Village, Ningcheng County, Inner Mongolia, China; Jiulongshan Formation, Middle Jurassic. Description. Relatively large insect, long 12.5 mm. The specimen shows a lateral view of the whole insect. The four wings folded together over the abdomen, showing only faint traces of the veins. Only venation of forewing is discernible from the four overlapped wings. Vertex of the head raised, the head capsule dongate, together with the dypeus, labrum and maxillae forms the prolonged rostrum, relatively slender and long. Compound eyes well developed. Antennae filiform, 11 segments visible, with short, stout scape and long slender tlageUum. Pronotum small, mesoand metanotum distinctly developed. Legs slender and long, covered with dense and tiny hairs, irregular arrangement. Coxae very dear, which are divided distincdy into eucoxae and mera. Trochanters small. Hind tibia obviously longer than femora, 1 terminal calcar visible. Tarsus 5-segnented, basitarsus longer than others. Forewing considerably large, length 12 mm, width 4 ram, apex rounded. Costal space not expanded. Pterosfigma dearly present. Sc long, 2-branched, forking nearly on a level with the origin of Rs, the upper branch short, almost uansverse, the lower long, extending to the pterosfigmal region, subcostal space very narrow. R1 simple, not branched, the wing margin, sigmoidally curved. Rs 4-branched, dividing into its two main branches nearly midwing, which forks nearly on the same level with M, R4 ＋ 5 forking on a level with R2 ＋ 3. M 4-branched, M3＋4 forking before M1＋2, which forks before the forking of R4＋ 5, the space between Rs and M very wide. CuA coalesced with M for a short distance basally, Cubito-median Y-vein present, having longer arm to CuA than to M, CuP simple, well separated from CuA. 1A, 2A simple, 2 cross-veins between them present. 3A simple. The veins of hind wing not discernible. Abdomen somewhat spindle-shaped, with 9 apparent segments, the 10th segment probably fused to the 9th segment, 1-segmented cersus visible, the termination of the cersus with a duster of distinct trichobothria. Gonocoxites evident, gonostyle tapering, visible a pair of paramenes. Netropanorpodes sentosus gen gen. et sp. nov. （Figs. 9-12, 15） Etymology. Sentosus from Latin, means many setae on the legs. Holotype. A well preserved female insect. Registration No： CNU-M-NN-2006026. Locality and horizon. Daohugou Village, N＇mgcheng County, Inner Mongolia, China; Jiulongshan Formation, Middle Jurassic. Description. Medium-sized insect, length 11 mm. The specimen shows a lateral view of the whole insect. The left forewing extending dorsal to the body, the right hind wing somewhat overlapping the body. The veins in the anal area are not discernible. The other two wings folded together, the veins are not discernible. Vertex of the head raised, rostrum slender and long, maxillary palp 4-segmented. Eyes large. Antennae 17 segments visible, with short, stout scape and long slender flagellum on which segmentation is evident. Pro- and mesonotum slightly larger, metanotum small. Legs slender and long, covered with dense and tiny hairs, irregular arrangement. Fore and middle legs with dear coxae, which are divided distinctly into eucoxae and mera, with slender femur slightly as long as tibiae. Trochanters small. Hind leg with coxa not discernible, tibia obviously longer than femora, with 2 terminal calcars, about equal in length. Tarsus 4 segments visible, Basitarsus longer than others, about two times as long as the second tarsomere. Forewing relatively broad, length 11.5 mm, width 4.5 mm, with rounded apical margin. Costal space expanded. Pterostigma dearly present, extending well below R1. Sc long, 2-branched, upper branch short, the lower one extending to the pterostigmal region. R1 simple, not branched, reaching the wing margin, sigmoidally curved near its termination. Rs 4-branched, dividing intO twO main branches before midwing, R4＋5 forking well before R2 ＋ 3, which almost forking on the same level with M3＋4. M arising from the base of the wing, 4-branched, forking on the same level with Rs, M3＋4 forking before M1 ＋2. M1 ＋2 forking after R4＋ 5, which forks before R2＋3. CuA coalesced with M for a long distance basaUy, cubito-median Y-vein evident. CuP and A not discernible. The hind wing smaller than forewing, but slightly broader. Sc a little shorter, only one branch. The veins of Cu and A not discernible. Other veins are similar to the forewing. Abdomen elongate, tapering apicaUy, with 11 dearly defined apparent segments. Each of the 9th to 11 th segments is telescoped into the preceding segment, the cercus not discernible. Comparison. The new species differs from Netropanorpodes decorosus sp. nov. in the following features on the forewing： the upper branch of Sc evident, R4＋ 5 forking well before R2 ＋ 3, which almost forking on the same level with M3 ＋ 4.
出处
《动物分类学报》
CSCD
北大核心
2007年第4期865-874,共10页
Acta Zootaxonomica Sinica
基金
国家自然科学基金(30370184
30430100)
北京市自然科学基金(5032003)
北京市教育委员会科技发展计划重点项目(KZ200410028013)
北京市市属市管高等学校拔尖创新人才项目共同资助.
关键词
长翅目
中生蝎蛉科
新属
新种
道虎沟
中侏罗世
Mecoptera,Mesopanorpodidae,new genus,new species,Daohugou,Middle Jurassic
