Twenty-three ichnotaxa have been found in the Silurian and Carboniferous turbidites of the Tianshan orogenic belt and the Triassic turbidites of the East Kunlun-West Qinling orogenic belt of northwestern China. They a...Twenty-three ichnotaxa have been found in the Silurian and Carboniferous turbidites of the Tianshan orogenic belt and the Triassic turbidites of the East Kunlun-West Qinling orogenic belt of northwestern China. They are Acanthorhaphe isp., ?Arthrophycus isp., Aulichnites parkerensis, Chondrites isp., C. intricatus, C. targionii, Cochlichnus anguineus, coprolite, Dendrotichnium haentzscheli, Helminthopsis isp., Helminthopsis abeli, H. hieroglyphica, H. cf. irregularis, Imbrichnus isp., Kunlunichnus qinghaiensis, Laevicyclus rotaeformis, Lophoctenium tianshanensis, Megagrapton isp., Micatuba verso?, Muensteria isp., Neonereites, Palaeophycus, and Zoophycos caudagalli, two of which, namely, Kunlunichnus qinghaiensis and Lophoctenium tianshanensis, are new ichnospecies. The described trace fossils can be grouped into five ichno-assemblages: the Aulichnites-Imbrichnus ichno-assemblage representing turbidity current deposits of a restricted anoxic deep-sea basin, the Lophoctenium ichno-assemblage showing the deep-sea or ocean environments relevant to a plate subduction, the Zoophycos-Helminthopsis ichnoassemblage representing the upper-middle turbidity fan deposits of a pelagic and hemipelagic environment, the Megagrapton-Chondrites ichno-assemblage representing the middle-lower turbidity fan deposits of an oceanic archipelago and the Kunlunichnus ichno-assemblage indicating bathyal turbidity current deposits.展开更多
Based on the analysis of trace fossils collected from the typical outcrop of the Lower Cambrian Wusongger Formation in the Kalpin area, ten ichnospecies of six ichnogenus were identified in the upper member of the Wus...Based on the analysis of trace fossils collected from the typical outcrop of the Lower Cambrian Wusongger Formation in the Kalpin area, ten ichnospecies of six ichnogenus were identified in the upper member of the Wusongger Formation, with most of them being found for the first time. The trace fossils are described seriatim. The ichnospecies are mainly represented by such common trace fossils as Ophiomorpha nodosa, Helminthopsis hieroglyphica, Helminthopsis ichnosp., Planolites beverleyensis, Planolites vulgaris, Planolites montanus, Palaeophycus striatus, Palaeophycus curvatus, Cochlichnus anguineus and Rituichnus elongatum. The trace fossils are of high diversity and low abundance, and can be diagnosed as the Cruziana ichnofacies. They can be interpreted as having formed in a shallow water environment.展开更多
The Luoping Biota discovered from the early Middle Triassic （Anisian） Guanling Formation of southwestern China represents a fully recovered shallow marine ecosystem, marking the end point of Early Triassic biotic re...The Luoping Biota discovered from the early Middle Triassic （Anisian） Guanling Formation of southwestern China represents a fully recovered shallow marine ecosystem, marking the end point of Early Triassic biotic recovery following the end-Permian mass extinction. Contemporaneously preserved are prolific trace fossils, which offer good opportunities to understand the palaeoecology of marine invertebrates from a fully recovered shallow marine ecosystem. Here we present a newly discovered sinuous branching burrow from the fossil-bearing unit in Member II of the Guanling Formation. Several features, including the horizontal regular sinuous nature, the branching pattern, typical H-junction, and the small wavelength/amplitude ratio of these sinuous structures within the burrow systems justify assignment of these traces as Sinusichnus sinuosus, a trace possibly produced by decapod crustaceans. Close association ofS. sinuosus with Rhizocorallium commune suggests a deposit-feeding strategy of these trace makers. The newly reported Anisian material from the Guanling Formation in Luoping represents first report of Sinusichnus from South China. The global record of Sinusichnus occurrence suggests that these burrows might have an older history than Early Middle Triassic.展开更多
There are abundant and highly varied trace fossils in the Sinian-Cambrian boundary beds in eastern Yunnan. Collections from 27 sections and their study give altogether 45 ichnogenera and 87 ichnospecies. Among them, 2...There are abundant and highly varied trace fossils in the Sinian-Cambrian boundary beds in eastern Yunnan. Collections from 27 sections and their study give altogether 45 ichnogenera and 87 ichnospecies. Among them, 2 ichnogenera and 7 ichnospecies are new, and 31 known ichnogenera and 38 known ichnospecies first found in the region. According to their order of appearance in the boundary beds and their assemblage features, the trace fossils in the Sinian-Cambrian boundary beds in the region may fall into 4 trace fossil assemblage zones in ascending order. In the paper they are correlated with those in the equivalent horizons abroad and their geological significance is discussed preliminarily.展开更多
Continental ichnology has been taking an ever increasing importance in stratigraphy. In this paper, Continental Trace Fossils in Semnan area have been studied. Semnan Province is one of the 31 provinces which is locat...Continental ichnology has been taking an ever increasing importance in stratigraphy. In this paper, Continental Trace Fossils in Semnan area have been studied. Semnan Province is one of the 31 provinces which is located in the north of Iran. In the East of Semnan City, Quaternary deposits occupy about 93% of the surface area, of which the Alluvial represents one of the most important geomorphological features in the area. Nevertheless, the outspread ichnological studies in Semnan City have received lesser attention in Continental Trace Fossils. Based on the systematic ichnology, 6 trace fossils are distinguished, namely Celliforma isp, Coprinisphaera isp, Palaeophycus tubularis, planolithes isp, Tombownichnus plenus, and Skolithos isp. The existence of Rhizoliths along with the tetrapod footprints and the survey of lithofacies all indicate that the trace fossils are related to Coprinisphaera ichnofacies. Based on ichnological and sedimentary facies analysis, the Continental Trace Fossils, in the study area, are located in the flood plain, point bar, and playa environment.展开更多
Six ichnogenera,Chondrites,Palaeophycus,Planolites,Rhizocorallium,Skolithos,and Thalassinoides,were identified from the drilling cores near the Global Stratotype Section and Point(GSSP) of Permian-Triassic boundary at...Six ichnogenera,Chondrites,Palaeophycus,Planolites,Rhizocorallium,Skolithos,and Thalassinoides,were identified from the drilling cores near the Global Stratotype Section and Point(GSSP) of Permian-Triassic boundary at Meishan,Zhejiang Province.The ichnological indices,including abundance,bioturbation index,and disturbed depth of trace fossils,show two episodes of intense changes near the Permian-Triassic Boundary(PTB).Episode I occurred in Beds 25-27b when the ecologically complicate forms such as Chondrites,Skolithos,Rhizocorallium,and Thalassinoides disappeared hereafter,the bioturbation index reduced from 1-5 to 1-3,and the disturbed depth declined from 5-66 to 2-5 cm.Episode II took place at the base of Bed 33 with the disappearance of Palaeophycus and Planolites,and subsequent absence of trace fossils and bioturbation till the middle-upper part of Bed 41 when the disturbed structures reoccurred,but they are only tiny Planolites and the bioturbation index was never higher than 3 and the disturbed depth less than 4 mm.Episode I shows an intense change,corresponding to the main stage of the end-Permian mass extinction,whereas Episode II is relatively weak,corresponding to the epilogue of the mass extinction of trace makers in the Early Triassic.Subsequently,ichnofossils were dominated by surface tracks in simple ecological habit and structures.This phenomenon indicates that the Early Triassic benthonic fauna is changed from sessile benthic system to mobile benthic system after the end-Permian mass extinction.In other words,the evolution of the trace fossils across the Permian-Triassic transition had an episodic process similar to the body fossils.In addition,the change of ichnofabrics is well coincided with the negative excursion of carbon isotopes and the expansion of cyanobacteria.As the results of physical and biogenic processes,trace fossils provided unique materials for the study of the biotic and environmental events,as well as their coupling evolution through the great Permian-Triassic transition.展开更多
The uppermost part of the Upper Bathonian Sponge Limestone member, Patcham Formation, of the Jhura Dome of Kachchh Mainland is a thickening-and shallowing-upward succession topped by medium-to thick-bedded hummocky cr...The uppermost part of the Upper Bathonian Sponge Limestone member, Patcham Formation, of the Jhura Dome of Kachchh Mainland is a thickening-and shallowing-upward succession topped by medium-to thick-bedded hummocky cross-stratified grainstones deposited by storm waves. Occasionally, thin, commonly lenticular, intraclastic-bioclastic silty marl intercalations between the grainstones are highly bioturbated, in contrast to the grainstones, in which, for the most part, trace fossils occur scattered. Large exposures of bedding planes of the grainstones allow the detailed investigation of ichnological features, whereas the high density of traces in the soft marls precludes the identification of any ichnotaxa. Eighteen ichnotaxa have been recorded including Ophiomorpha, Thalassinoides, Taenidium, Gyrophyllites, Chondrites, Dactyloidites, Teichichnus, Bolonia, and Ancorichnus. Except for Ophiomorpha nodosa and Thalassinoides, which generally indicate moderate to high energy conditions and are the dwelling burrows of suspension-feeding to omnivorous crustaceans, the ichnotaxa represent a deposit-feeding behaviour of their producers and thus are characteristic of low-energy environments. The trace fossils form three ichnoassemblages characterized by(1) Ophiomorpha nodosa and Thalassinoides suevicus,(2)?Thalassinoides isp. A, Taenidium, and Bolonia lata,and(3) Ancorichnus. The dominance of traces of deposit-feeders in rocks indicative of high-energy events is counterintuitive and points to their non-contemporaneity. The sediments were deposited during brief highenergy events, whereas the trace fossils were produced when, after waning of storms low-energy conditions prevailed. This time-averaging is particularly pronounced in trace fossils that extend vertically downwards and may reach strata deposited under distinctly different conditions. Thus, environmental interpretations based on trace fossils should refer to colonisation surfaces rather than to the sediment surrounding the trace fossils. In the latter case, interpretations may be erroneous, especially when erosion subsequently destroyed sedimentary evidence of their original environment.展开更多
THE origin and evolution of Precambrian lives has been a long-standing controversial problem. It is now generally accepted by most paleontologists that the origin of metazoa can be traced back to more than 1000 Ma, be...THE origin and evolution of Precambrian lives has been a long-standing controversial problem. It is now generally accepted by most paleontologists that the origin of metazoa can be traced back to more than 1000 Ma, because the Ediacara and latest pre-Ediacara fauna, such as the Huainan fauna, are featured by their diversification of forms, complicated structures展开更多
Extremely abundant trace fossils are associated with the deep-sea deposits, especially in turbidite measure and flysch facies. And these trace groups then become the effective indicators to reconstruct the ancient dep...Extremely abundant trace fossils are associated with the deep-sea deposits, especially in turbidite measure and flysch facies. And these trace groups then become the effective indicators to reconstruct the ancient depositional environment and the palaeogeography. This paper focuses on ( ⅰ ) the formational features of deep-sea fossil and its assemblage feature, and further recongnizes that the coexistence of the shallow and deep water trace group in a depositional sequences is the identification mark of turbidite sequence; ( ⅱ ) thought and methods using the trace fossil to restore their environmental parameters such as the ancient sea depth, oxygenbearing condition and sedimentation rate, etc.; ( ⅲ ) the application and significance of deep-sea ichnofacies in environment interpretation.展开更多
A new taxonomy on trace fossils considering both the morphological and structural features is suggested. From mathematical and paleobiological viewpoints, all trace fossils can be ordered into five classes of the hier...A new taxonomy on trace fossils considering both the morphological and structural features is suggested. From mathematical and paleobiological viewpoints, all trace fossils can be ordered into five classes of the hierarchy: topological ichnocategory, ichnofamily, ichnosubfamily, ichnogenus and ichnospecies. It is hoped that by means of this taxonomy, the effects such as tectonic deformation can no longer influence the fossils’ morphological analysis and the configuration complexities within various trace fossils can also be compared to one another immediately.展开更多
文摘Twenty-three ichnotaxa have been found in the Silurian and Carboniferous turbidites of the Tianshan orogenic belt and the Triassic turbidites of the East Kunlun-West Qinling orogenic belt of northwestern China. They are Acanthorhaphe isp., ?Arthrophycus isp., Aulichnites parkerensis, Chondrites isp., C. intricatus, C. targionii, Cochlichnus anguineus, coprolite, Dendrotichnium haentzscheli, Helminthopsis isp., Helminthopsis abeli, H. hieroglyphica, H. cf. irregularis, Imbrichnus isp., Kunlunichnus qinghaiensis, Laevicyclus rotaeformis, Lophoctenium tianshanensis, Megagrapton isp., Micatuba verso?, Muensteria isp., Neonereites, Palaeophycus, and Zoophycos caudagalli, two of which, namely, Kunlunichnus qinghaiensis and Lophoctenium tianshanensis, are new ichnospecies. The described trace fossils can be grouped into five ichno-assemblages: the Aulichnites-Imbrichnus ichno-assemblage representing turbidity current deposits of a restricted anoxic deep-sea basin, the Lophoctenium ichno-assemblage showing the deep-sea or ocean environments relevant to a plate subduction, the Zoophycos-Helminthopsis ichnoassemblage representing the upper-middle turbidity fan deposits of a pelagic and hemipelagic environment, the Megagrapton-Chondrites ichno-assemblage representing the middle-lower turbidity fan deposits of an oceanic archipelago and the Kunlunichnus ichno-assemblage indicating bathyal turbidity current deposits.
基金supported by National Natural Science Foundation of China (No. 41072102)the National Basic Research Program of China(No.2012CB214804)
文摘Based on the analysis of trace fossils collected from the typical outcrop of the Lower Cambrian Wusongger Formation in the Kalpin area, ten ichnospecies of six ichnogenus were identified in the upper member of the Wusongger Formation, with most of them being found for the first time. The trace fossils are described seriatim. The ichnospecies are mainly represented by such common trace fossils as Ophiomorpha nodosa, Helminthopsis hieroglyphica, Helminthopsis ichnosp., Planolites beverleyensis, Planolites vulgaris, Planolites montanus, Palaeophycus striatus, Palaeophycus curvatus, Cochlichnus anguineus and Rituichnus elongatum. The trace fossils are of high diversity and low abundance, and can be diagnosed as the Cruziana ichnofacies. They can be interpreted as having formed in a shallow water environment.
基金partly supported by the ARCDP discoverygrant to G. R. Shi (No. DP150100690)supported by a CAS Pioneer Hundred Talents Program (to ML)+2 种基金supported by a grant from the State Key Laboratory of Biogeology and Environmental Biology (No. GBL21410 to ML), China University of Geosciences (Wuhan)a grant from the National Natural Science Foundation of China (No. 41502013)supported by for China Geological Survey projects (Nos. DD20160020, 1212011140051, 12120114030601, and 1212010610211)
文摘The Luoping Biota discovered from the early Middle Triassic （Anisian） Guanling Formation of southwestern China represents a fully recovered shallow marine ecosystem, marking the end point of Early Triassic biotic recovery following the end-Permian mass extinction. Contemporaneously preserved are prolific trace fossils, which offer good opportunities to understand the palaeoecology of marine invertebrates from a fully recovered shallow marine ecosystem. Here we present a newly discovered sinuous branching burrow from the fossil-bearing unit in Member II of the Guanling Formation. Several features, including the horizontal regular sinuous nature, the branching pattern, typical H-junction, and the small wavelength/amplitude ratio of these sinuous structures within the burrow systems justify assignment of these traces as Sinusichnus sinuosus, a trace possibly produced by decapod crustaceans. Close association ofS. sinuosus with Rhizocorallium commune suggests a deposit-feeding strategy of these trace makers. The newly reported Anisian material from the Guanling Formation in Luoping represents first report of Sinusichnus from South China. The global record of Sinusichnus occurrence suggests that these burrows might have an older history than Early Middle Triassic.
基金This research was a project (No. 4870087) supported by the National Natural Science Foundation of China.
文摘There are abundant and highly varied trace fossils in the Sinian-Cambrian boundary beds in eastern Yunnan. Collections from 27 sections and their study give altogether 45 ichnogenera and 87 ichnospecies. Among them, 2 ichnogenera and 7 ichnospecies are new, and 31 known ichnogenera and 38 known ichnospecies first found in the region. According to their order of appearance in the boundary beds and their assemblage features, the trace fossils in the Sinian-Cambrian boundary beds in the region may fall into 4 trace fossil assemblage zones in ascending order. In the paper they are correlated with those in the equivalent horizons abroad and their geological significance is discussed preliminarily.
文摘Continental ichnology has been taking an ever increasing importance in stratigraphy. In this paper, Continental Trace Fossils in Semnan area have been studied. Semnan Province is one of the 31 provinces which is located in the north of Iran. In the East of Semnan City, Quaternary deposits occupy about 93% of the surface area, of which the Alluvial represents one of the most important geomorphological features in the area. Nevertheless, the outspread ichnological studies in Semnan City have received lesser attention in Continental Trace Fossils. Based on the systematic ichnology, 6 trace fossils are distinguished, namely Celliforma isp, Coprinisphaera isp, Palaeophycus tubularis, planolithes isp, Tombownichnus plenus, and Skolithos isp. The existence of Rhizoliths along with the tetrapod footprints and the survey of lithofacies all indicate that the trace fossils are related to Coprinisphaera ichnofacies. Based on ichnological and sedimentary facies analysis, the Continental Trace Fossils, in the study area, are located in the flood plain, point bar, and playa environment.
基金supported by National Basic Research Program of China (Grant No. 2006CB80640)National Natural Science Foundation of China (Grant Nos. 40830212,40772022)
文摘Six ichnogenera,Chondrites,Palaeophycus,Planolites,Rhizocorallium,Skolithos,and Thalassinoides,were identified from the drilling cores near the Global Stratotype Section and Point(GSSP) of Permian-Triassic boundary at Meishan,Zhejiang Province.The ichnological indices,including abundance,bioturbation index,and disturbed depth of trace fossils,show two episodes of intense changes near the Permian-Triassic Boundary(PTB).Episode I occurred in Beds 25-27b when the ecologically complicate forms such as Chondrites,Skolithos,Rhizocorallium,and Thalassinoides disappeared hereafter,the bioturbation index reduced from 1-5 to 1-3,and the disturbed depth declined from 5-66 to 2-5 cm.Episode II took place at the base of Bed 33 with the disappearance of Palaeophycus and Planolites,and subsequent absence of trace fossils and bioturbation till the middle-upper part of Bed 41 when the disturbed structures reoccurred,but they are only tiny Planolites and the bioturbation index was never higher than 3 and the disturbed depth less than 4 mm.Episode I shows an intense change,corresponding to the main stage of the end-Permian mass extinction,whereas Episode II is relatively weak,corresponding to the epilogue of the mass extinction of trace makers in the Early Triassic.Subsequently,ichnofossils were dominated by surface tracks in simple ecological habit and structures.This phenomenon indicates that the Early Triassic benthonic fauna is changed from sessile benthic system to mobile benthic system after the end-Permian mass extinction.In other words,the evolution of the trace fossils across the Permian-Triassic transition had an episodic process similar to the body fossils.In addition,the change of ichnofabrics is well coincided with the negative excursion of carbon isotopes and the expansion of cyanobacteria.As the results of physical and biogenic processes,trace fossils provided unique materials for the study of the biotic and environmental events,as well as their coupling evolution through the great Permian-Triassic transition.
基金supported by the Jagiellonian University(DS funds)financial assistance for the fieldwork by DST(project EMR/ 2015/001574)financial support by the Alexander von Humboldt Foundation
文摘The uppermost part of the Upper Bathonian Sponge Limestone member, Patcham Formation, of the Jhura Dome of Kachchh Mainland is a thickening-and shallowing-upward succession topped by medium-to thick-bedded hummocky cross-stratified grainstones deposited by storm waves. Occasionally, thin, commonly lenticular, intraclastic-bioclastic silty marl intercalations between the grainstones are highly bioturbated, in contrast to the grainstones, in which, for the most part, trace fossils occur scattered. Large exposures of bedding planes of the grainstones allow the detailed investigation of ichnological features, whereas the high density of traces in the soft marls precludes the identification of any ichnotaxa. Eighteen ichnotaxa have been recorded including Ophiomorpha, Thalassinoides, Taenidium, Gyrophyllites, Chondrites, Dactyloidites, Teichichnus, Bolonia, and Ancorichnus. Except for Ophiomorpha nodosa and Thalassinoides, which generally indicate moderate to high energy conditions and are the dwelling burrows of suspension-feeding to omnivorous crustaceans, the ichnotaxa represent a deposit-feeding behaviour of their producers and thus are characteristic of low-energy environments. The trace fossils form three ichnoassemblages characterized by(1) Ophiomorpha nodosa and Thalassinoides suevicus,(2)?Thalassinoides isp. A, Taenidium, and Bolonia lata,and(3) Ancorichnus. The dominance of traces of deposit-feeders in rocks indicative of high-energy events is counterintuitive and points to their non-contemporaneity. The sediments were deposited during brief highenergy events, whereas the trace fossils were produced when, after waning of storms low-energy conditions prevailed. This time-averaging is particularly pronounced in trace fossils that extend vertically downwards and may reach strata deposited under distinctly different conditions. Thus, environmental interpretations based on trace fossils should refer to colonisation surfaces rather than to the sediment surrounding the trace fossils. In the latter case, interpretations may be erroneous, especially when erosion subsequently destroyed sedimentary evidence of their original environment.
文摘THE origin and evolution of Precambrian lives has been a long-standing controversial problem. It is now generally accepted by most paleontologists that the origin of metazoa can be traced back to more than 1000 Ma, because the Ediacara and latest pre-Ediacara fauna, such as the Huainan fauna, are featured by their diversification of forms, complicated structures
文摘Extremely abundant trace fossils are associated with the deep-sea deposits, especially in turbidite measure and flysch facies. And these trace groups then become the effective indicators to reconstruct the ancient depositional environment and the palaeogeography. This paper focuses on ( ⅰ ) the formational features of deep-sea fossil and its assemblage feature, and further recongnizes that the coexistence of the shallow and deep water trace group in a depositional sequences is the identification mark of turbidite sequence; ( ⅱ ) thought and methods using the trace fossil to restore their environmental parameters such as the ancient sea depth, oxygenbearing condition and sedimentation rate, etc.; ( ⅲ ) the application and significance of deep-sea ichnofacies in environment interpretation.
文摘A new taxonomy on trace fossils considering both the morphological and structural features is suggested. From mathematical and paleobiological viewpoints, all trace fossils can be ordered into five classes of the hierarchy: topological ichnocategory, ichnofamily, ichnosubfamily, ichnogenus and ichnospecies. It is hoped that by means of this taxonomy, the effects such as tectonic deformation can no longer influence the fossils’ morphological analysis and the configuration complexities within various trace fossils can also be compared to one another immediately.
