Larger benthic foraminifera during the Eocene within the Tethyan realm played a crucial role in regional biostratigraphy.In the Middle East,particularly in Libya,Tunisia,and Egypt,nummulitids are key constituents of n...Larger benthic foraminifera during the Eocene within the Tethyan realm played a crucial role in regional biostratigraphy.In the Middle East,particularly in Libya,Tunisia,and Egypt,nummulitids are key constituents of nummulitic limestone reservoirs,making the taxonomic identification of this structurally complex group vital for biostratigraphic and petroleum applications.This review focuses on the genus Gaziryina of the nummulitids,clarifying its taxonomic status,biostratigraphic utility,and paleobiogeographic significance using biometric data from published literature.Previously misidentified as Nummulites pulchellus,Gaziryina species have been systematically reassessed based on biometric data from published literature.The findings confirm Gaziryina as a distinct genus comprising two species:Gaziryina basatinensis(late Lutetian-Bartonian,SBZ15-18a) and Gaziryina pulchellus(late Bartonian-Priabonian,SBZ18a/b-20).Morphological characters(test shape,size progression,chamber configuration,protoconch enlargement) and ontogenetic trends(tight to lax) indicate a phylogenetic transition where Gaziryina basatinensis evolved into Gaziryina pulchellus,consistent with Cope's rule.Paleobiogeographic reconstruction indicates a probable origin in the southern NeoTethys,likely the Western Desert of Egypt,with subsequent dispersal across the northern and southeastern Tethys.These findings confirm Gaziryina as a key biostratigraphic marker,necessitating further research on its evolution and biostratigraphic framework.展开更多
A comprehensive compilation and systematic analysis of known early and middle Permian brachiopod faunas shows that the early Permian brachiopod faunas comprise three realms,six regions,and eleven provinces,while those...A comprehensive compilation and systematic analysis of known early and middle Permian brachiopod faunas shows that the early Permian brachiopod faunas comprise three realms,six regions,and eleven provinces,while those of the middle Permian comprise three realms,four regions,and eight provinces.A comparison and analysis of brachiopod faunal patterns reveal a coevolution between global brachiopod paleobiogeography and tectonopaleogeography during the early-middle Permian.Although temperature/latitude is the main factor controlling the formation of three realms,tectonopaleogeographic factors determine the temperature/latitude in which the continents were located.The'continental barrier'of Pangea,as a'central axis'continent,divided the three realms into six regions,which indicates that the formation of biogeographic regions was controlled mainly by the tectonopaleogeographic factors.The evolution of tectonopaleogeography was sometimes a long-term process,so that the biogeographic regions(or provinces)controlled by tectonopaleogeography displayed relative stability.Shifts in the nature of biogeographic provinces(e.g.,from cool water to warm water,and vice versa),extensions or narrowing of geographical ranges,and recombinations of some provinces were all related to regional tectonic evolution.The study of the coevolution between brachiopod paleobiogeography and tectonopaleogeography not only accounts for the formation mechanisms of brachiopod paleobiogeographic patterns during the early-middle Permian,but also provides evidences for the locations and configurations of oceans and plates(blocks)during this period.展开更多
Although the fossil record of non-avian dinosaurs from the Cretaceous of Antarctica is the poorest of any continent,fossils representing at least five major taxonomic groups(Ankylosauria,early-diverging Ornithopoda,Ha...Although the fossil record of non-avian dinosaurs from the Cretaceous of Antarctica is the poorest of any continent,fossils representing at least five major taxonomic groups(Ankylosauria,early-diverging Ornithopoda,Hadrosauridae,Titanosauria,and Theropoda)have been recovered.All come from Upper Cretaceous(Coniacian–Maastrichtian)marine and nearshore deposits belonging to the Gustav and Marambio groups of the James Ross Basin at the northern tip of the Antarctic Peninsula.The majority of these finds have come from the Campanian–Maastrichtian Snow Hill Island and López de Bertodano formations of James Ross and Vega islands.Given the rarity of Antarctic Cretaceous non-avian dinosaurs,discoveries of any fossils of these archosaurs,no matter how meager,are of significance.Here we describe fragmentary new ornithischian(ankylosaur and ornithopod)material from the upper Campanian–lower Maastrichtian Cape Lamb Member of the Snow Hill Island Formation and the Maastrichtian Sandwich Bluff Member of the López de Bertodano Formation.One of these specimens is considered to probably pertain to the holotypic individual of the early-diverging ornithopod Morrosaurus antarcticus.We also provide an up-to-date synthesis of the Late Cretaceous non-avian dinosaur record of the James Ross Basin and analyze the biostratigraphic occurrences of the various finds,demonstrating that most(including all named taxa and all reasonably complete skeletons discovered to date)occur within a relatively condensed temporal interval of the late Campanian to early Maastrichtian.Most or all James Ross Basin dinosaurs share close affinities with penecontemporaneous taxa from Patagonia,indicating that at least some continental vertebrates could disperse between southern South America and Antarctica during the final stages of the Mesozoic.展开更多
New well-preserved rudist materials come from the Barremian–Turonian marine formations in the Yelleg, Minsherah, Maaza, and Raghawi sections in North Sinai, northern Egypt. There, 17 discovered rudist species belongi...New well-preserved rudist materials come from the Barremian–Turonian marine formations in the Yelleg, Minsherah, Maaza, and Raghawi sections in North Sinai, northern Egypt. There, 17 discovered rudist species belonging to 13 genera and seven taxonomic families are described in detail as follows: Eoradiolites plicatus(Conrad), Eoradiolites liratus(Conrad), Praeradiolites ponsianus(d'Archiac), Archaeoradiolites sp., Bournonia africana Douvillé, Bournonia fourtaui Douville, Biradiolites lombricalis(d'Orbigny), Biradiolites zumoffeni Douville, Radiolites lewyi lewyi Parnes, Radiolites sauvagesi(d'Hombres-Firmas), Durania arnaudi(Choffat), Toucasia carinata(Matheron), Toucasia sp., Neocaprina raghawiensis Steuber and Bachmann, Sellaea sp., Ichthyosarcolites sp. and Horoiopleura sp.. The Cenomanian deposits in the northern Eastern desert of Egypt only contain E. liratus. The domination of the Cenomanian rudist species in North Sinai, however, is attributed to changes in the platform, which passes mainly from carbonate in the north to siliciclastics in the south. The presence of Horiopleura sp. in the late Barremian-early Aptian deposits is documented for the first time, which suggests the expansion of taxa of the genus Horiopleura Douville to North Sinai during this interval. The age of the rudist species is documented from the late Barremian-Turonian formations, with their geographic distribution in the Mediterranean region considered..展开更多
The distribution of ostracods and benthonic foraminifers in the China sea area is briefly reviewed from the paleobio-geographic viewpoint in this paper. Three regions can be distinguished in the area on the basis of m...The distribution of ostracods and benthonic foraminifers in the China sea area is briefly reviewed from the paleobio-geographic viewpoint in this paper. Three regions can be distinguished in the area on the basis of modern distribution data: Region I (the Huanghai Sea and the Bohai Sea) with cool and temperate forms, Region I (the East China Sea and the northern part of the South China Sea) with subtropical warm-water forms and Region Ⅲ (central and southern parts of the South China Sea) with larger foraminifers and other tropical warm-water forms. The occurrence of Nummulites-Discocyclira fauna in the Eocene deposits of the East China Sea indicates a northward extension of tropical zoogeographical region at the time, whereas the distribution pattern of the Miocene Nephrolepidina-Miogypsiua-Austrotrillina fauna in the South China Sea resembles that of the present larger-foraminiferal fauna. In the South China Sea and Taiwan, a warm-water fauna with Asterorotalia and Pseudorotalia first appeared in late Miocene and then flourished in the Pliocene. Its northward invasion into the Bohai Sea area during late Pleistocene may be related to changes in current system. Many of the endemic foraminiferal and ostracod genera and species ( Sinocytheridea etc. ) in recent coastal faunas probably have originated from the South China Sea, and their occurrence in Japan or Australia may be indicative of ties between these areas during glacial times.展开更多
Porcupines,members of the Hystricidae family,represent a unique group of herbivorous mammals.This study details the identification of a newly discovered mandible fragment of Hystrix primigenia,along with a right cheek ...Porcupines,members of the Hystricidae family,represent a unique group of herbivorous mammals.This study details the identification of a newly discovered mandible fragment of Hystrix primigenia,along with a right cheek tooth series from the middle Turolian Kemiklitepe-A fossil locality.While Hystrix fossils are found in numer-ous localities,the materials are often limited to a few dental fragments or isolated teeth,posing challenges to systematic investigations.The examination of this lower tooth series prompted a comprehensive review of all H.primigeniafindings across Eurasia,shedding light on its adaptive characteristics over time and space.Our paleo-biogeographical analysis indicates the absence of H.depereti in Late Miocene Türkiye,while the dispersal range of H.primigenia is broader than that of H.depereti in Eurasia.Additionally,the study delves into the discussion of H.primigenia and H.deperetifindings in Eurasia,ultimately refining the categorization of Late Miocene Hys-trix discoveries in Türkiye to two species:H.primigenia and H.kayae.Our review suggests the possibility of an additional H.kayaefinding from Samos,Greece.展开更多
The Neogene interval witnessed three major waves of intercontinental dispersals by carnivoran mammals,mainly migrating from Eurasia to North America but with rare occurrences going in the opposite direction(e.g.,Sthen...The Neogene interval witnessed three major waves of intercontinental dispersals by carnivoran mammals,mainly migrating from Eurasia to North America but with rare occurrences going in the opposite direction(e.g.,Sthenictis,Leptarctus).The timing and composition of dispersers are thought to be controlled by filter-bridge mechanisms,but it is unclear what types of regional conditions promoted occasional dispersals outside of the three major waves.We study and report on new occurrences of small-bodied carnivorans from late Clarendonian-aged Black Hawk Ranch and Cuyama Valley fossil mammal assemblages in California.The relatively late occurrences of the mustelids Sthenictis and Hoplictis and the ailurid Alopecocyon in coastal regions of western North America suggest that nearshore dispersal corridors both facilitate and preserve faunal elements later than they might otherwise occur further inland.The availability of both marine-and terrestrially-derived food resources may be an important factor in allowing coastally dispersing taxa to be accommodated in those ecological communities compared to less heterogenous environments further inland.展开更多
A new species of Bothriolepis(Placodermi,Antiarcha),B.zhujiangyuanensis sp.nov.,is described from the Eifelian(Middle Devonian)Shangshuanghe Formation at the Pearl River Source Scenic Area,Qujing,Yunnan Province,south...A new species of Bothriolepis(Placodermi,Antiarcha),B.zhujiangyuanensis sp.nov.,is described from the Eifelian(Middle Devonian)Shangshuanghe Formation at the Pearl River Source Scenic Area,Qujing,Yunnan Province,southwestern China and is characterized by a broad postpineal plate bearing a straight anterior margin,a premedian plate wider than long,and a weakly developed dorsal median ridge of the trunk shield.As the eighth Bothriolepis taxon in China,B.zhujiangyuanensis sp.nov.provides a wealth of morphological data to clarify the intrageneric interrelationships.Maximum parsimony analysis,using a revised data set of the genus with 76 ingroup taxa and 72 characters,yields a well-resolved consensus tree for Bothriolepis.The phylogenetic result shows a sister-group relationship between B.zhujiangyuanensis and B.babichevi from Kazakhstan,corroborating the biogeographic proximity between the South China and Kazakhstania blocks during the Middle Devonian.展开更多
Ten palm leaf impressions are documented from the latest Maastrichtian(late Cretaceous) to early Danian(earliest Paleocene) sediments(K-Pg,c.66-64 Ma) of the Mandla Lobe of the Deccan Intertrappean Beds,Madhya Pradesh...Ten palm leaf impressions are documented from the latest Maastrichtian(late Cretaceous) to early Danian(earliest Paleocene) sediments(K-Pg,c.66-64 Ma) of the Mandla Lobe of the Deccan Intertrappean Beds,Madhya Pradesh,central India.The palmate leaf shape along with a definite wellpreserved costa support their placement in the subfamily Coryphoideae of the family Arecaceae.We place all recovered palm leaf specimens in the fossil genus Sabalites,report seven species of coryphoid palms and describe two new species namely,Sabalities umariaensis sp.nov.and Sabalites ghughuaensis sp.nov.The fossils indicate that coryphoid palms were highly diverse in central India by the latest Cretaceous.These and earlier reported coryphoid palm fossils from the same locality indicate that they experienced a warm and humid tropical environment during the time of deposition.These discoveries confirm the presence of a diversity of Coryphoideae in Gondwana prior to the India-Eurasia collision and provide information about coryphoid biogeographical history over geological time.Based on megafossil remains,we trace coryphoid palm migration pathways from India to mainland Southeast(SE) Asia and other parts of Asia after the docking of the Indian subcontinent with Eurasia early in the Paleogene.展开更多
Ordovician conodonts representing 28 genera and 28 named and three unnamed species were identified from 740 chert and siliceous siltstone spot samples(>3000 thin sections)from deep-water turbiditic sequences of the...Ordovician conodonts representing 28 genera and 28 named and three unnamed species were identified from 740 chert and siliceous siltstone spot samples(>3000 thin sections)from deep-water turbiditic sequences of the Lachlan Orogen in central and southern New South Wales,Australia.Based on these faunas,a new conodont biozonal scheme has been established to divide the Ordovician turbiditic successions of the Lachlan Orogen into 12 superbiozones and biozones.They are(in ascending order)the Paracordylodus gracilis Superbiozone(including the Prioniodus oepiki Biozone),Periodon flabellum Superbiozone(including the Oepikodus evae Biozone in the lower part),Periodon hankensis Biozone,Periodon aculeatus Superbiozone(including the Histiodella labiosa,Histiodella holodentata,Histiodella kristinae,Pygodus serra and Pygodus anserinus biozones)and the Periodon grandis Biozone.The Pygodus anserinus Biozone is divided further into the lower and upper subbiozones.This new conodont biozonation scheme spanning the upper Tremadocian to middle Katian interval permits precise age-dating and correlation of deep-water siliciclastic rocks that characterize the Ordovician Deep-Sea Realm regionally and internationally.展开更多
Recent geological and paleontological exploration in the Indus basin of Pakistan allowed the discoveries of numerous remains of non-marine reptiles( titanosaurian sauropod,abelisaurian and noasaurian theropod dinosaur...Recent geological and paleontological exploration in the Indus basin of Pakistan allowed the discoveries of numerous remains of non-marine reptiles( titanosaurian sauropod,abelisaurian and noasaurian theropod dinosaurs),and marine reptiles( crocodiles),flying reptiles( pterosaurs),marine and non-marine mammals,fishes,invertebrates,and plants,especially Pakistan is relatively rich in footprints / trackways in the Mesozoic.These vertebrates of Indo-Pakistan are very significant for paleobiogeographic study due to the present-day connection of this continent with Asia in Northern Hemisphere,whereas during past( Jurassic and pre-Jurassic) it was connected to the Gondwana. The Mesozoic vertebrates show close affinities with Gondwanan landmasses.The Cenozoic vertebrates show Eurasian affinity and migrated from Indo-Pak subcontinent to Eurasia or vice versa via Paleo Indus River systems along Western Indus Suture,after long journey of about 6 000 km the first collision of Indo-Pak subcontinent with Asia occurred at terminal Cretaceous.展开更多
The first unequivocal remains of medium to large-sized mustelids from the middle Miocene Halamagai and Kekemaideng formations have been found in the Ulungur River area, Xinjiang, northwestern China. These new fossils ...The first unequivocal remains of medium to large-sized mustelids from the middle Miocene Halamagai and Kekemaideng formations have been found in the Ulungur River area, Xinjiang, northwestern China. These new fossils are referred to the hypercarnivorous mustelid Hoplictis Ginsburg, 1961 and denote the first record of the genus in East Asia.We define Hoplictis baihu n. sp., for the mustelid from Tieersihabahe(Halamagai Fm.), which represents the smallest species of the genus. This primitive form is closer to H. florancei and H. noueli than to H. anatolicus and later, larger and more derived Hoplictis spp., from Europe and North America. A large toothless mandible from Duolebulejin(Kekemaiden Fm.) is assigned to Hoplictis cf. helbingi, and it presumably might represent the first record of H. helbingi outside Western Europe. The systematic position of Hoplicitis in relation to Ischyrictis with which it is similar is clarified. The occurrence in East Asia of two species of Hoplictis greatly expands the known distribution and diversity of the genus, and supports a Palaearctic Neogene dispersal event of carnivorans between Europe and Asia during the late Shanwangian–early Tunggurian equivalent to MN5–6 in Europe, and indication of another dispersal event from Europe to North America,through Northwest China during the late Tunggurian, equivalent to MN7–8 in Europe.展开更多
On the basis of a worldwide review of Claraia , Pseudoclaraia and Eumorphotis , especially their important species , two range zones are recognized:the Pseudoclaraia wangi range zone —— Upper Griesbachian——and the...On the basis of a worldwide review of Claraia , Pseudoclaraia and Eumorphotis , especially their important species , two range zones are recognized:the Pseudoclaraia wangi range zone —— Upper Griesbachian——and the Claraia stachei- C. aurita - Eumorphotis multiformis acme zone——Upper Griesbachian to Lower Smithian This result revises both the tradition-al Chinese concept regarding these fossils as limited in Lower Scythian and the viewpoint of some Eurasian researchers who hold Eumorphotis multiformis to be Upper Scythian .The paleogeographical distribution of the three genera is controlled primarily by temperature (latitude) and secondarily by physiographic separations .Distribution of these fossils is subdivided into four realms and five provinces . The Eastern Asia province of the Tethys realm is suggested as the source area of these fossils .The striking contrast between the composition of the species along the southern and northern margins of the Tethys , together with other distributional characters ,supports the plate tectonic theory .展开更多
Numerous new records of Ferganiella, Podozamites, and Schidolepium, including a new species, Ferganiella ivantsovii sp. nov., are described from the Early Jurassic(Toarcian) Middle Subformation of the Prisayan Formati...Numerous new records of Ferganiella, Podozamites, and Schidolepium, including a new species, Ferganiella ivantsovii sp. nov., are described from the Early Jurassic(Toarcian) Middle Subformation of the Prisayan Formation from the Euro-Sinian paleofloristic region in the Irkutsk Basin, Eastern Siberia, Russia. An analysis of the paleogeographic distribution of Ferganiella and Podozamites shows that both genera were the most diverse and numerous in the East Asian province of the Euro-Sinian region and in the Northern Chinese province of the Siberian region during the Early and Middle Jurassic. These phytochoria were located in the subtropical and temperate subtropical climate zones, which allows us to consider Ferganiella and Podozamites as thermophilic plants, which are important indicators of the Early Toarcian climatic optimum. Their abundance in the Irkutsk Basin thus may indicate Early Toarcian warming;further abundant Schidolepium cones, which produced Araucariacites pollen, typical for Euro-Sinian flora complement the scenario. Thus, the new finds are the first macrofloristic indicators of the Toarcian climatic optimum in the Irkutsk Basin.展开更多
Geologic studies of the Late Cretaceous-Paleogene stratigraphic record of southern Tibet reveal distinct episodes of collision and post-collisional compression.The composition of sediments exposed along the Indus-Yarl...Geologic studies of the Late Cretaceous-Paleogene stratigraphic record of southern Tibet reveal distinct episodes of collision and post-collisional compression.The composition of sediments exposed along the Indus-Yarlung Zangbo suture,tectonic deformation,展开更多
Saraikistan (South Punjab and surrounding) area of Pakistan is located in the central Pakistan. This area represents Triassic-Jurassic to Recent sedimentary marine and terrestrial strata. Most of the Mesozoic and Earl...Saraikistan (South Punjab and surrounding) area of Pakistan is located in the central Pakistan. This area represents Triassic-Jurassic to Recent sedimentary marine and terrestrial strata. Most of the Mesozoic and Early Cenozoic are represented by marine strata with rare terrestrial deposits, while the Late Cenozoic is represented by continental fluvial deposits. This area hosts significant mineral deposits and their development can play a significant role in the development of Saraikistan region and ultimately for Pakistan. The data of recently discovered biotas from Cambrian to Miocene age are tabulated for quick view. Mesozoic biotas show a prominent paleobiogeographic link with Gondwana and Cenozoic show Eurasian. Phylogeny and hypodigm of Poripuchian titanosaurs from India and Pakistan are hinted at here.展开更多
The overall occurrence,stratigraphical distribution,palaeoenvironmental and palaeobiogeographical significances of the ostracod faunas from the Silurian Pulu Formation in the Yalai WestⅡsection,Nyalam region,southern...The overall occurrence,stratigraphical distribution,palaeoenvironmental and palaeobiogeographical significances of the ostracod faunas from the Silurian Pulu Formation in the Yalai WestⅡsection,Nyalam region,southern Tibet,China,are documented for the first time.Thirty-two species belonging to 19 genera are identified and figured herein.The ostracod fauna in the Pulu Formation indicates an age of the late Llandovery-Pridoli.The ostracods belong to a podocope-rich association,which suggests an offshore environment for the Yalai West II section.The ostracods of the Pulu Formation have strong affinities with the South China taxa and also share some similarities with those from Baltica.This implies that ostracods could probably benefit from sea-level changes to facilitate faunal exchanges between peri-Gondwana and Baltica.Geographical isolation and global sea-level changes are proposed as the primary factors controlling the palaeobiogeographical distribution of ostracods during the Silurian.展开更多
End-Triassic ammonoid and bivalve faunas of the Germig area, Tibetan Himalaya, lived in a tropical, shallow-water environment during the Triassic-Jurassic boundary interval. High stratigraphic resolution based on ammo...End-Triassic ammonoid and bivalve faunas of the Germig area, Tibetan Himalaya, lived in a tropical, shallow-water environment during the Triassic-Jurassic boundary interval. High stratigraphic resolution based on ammonite-biochrons allows to tracing the place of origin of several faunal elements. The bivalves Aguilerella and Ctenostreon occurred first in the Tibetan Himalaya and migrated from there to the eastern South Pacific, exhibiting a pantropic dispersal pattern. This dispersal route is supported by the distribution pattern of the ammonites Choristoceras, Discamphiceras, Pleuroacanthites, and Psiloceras calliphyllum. A few taxa, which went extinct everywhere else by the end of the Triassic, survived in the Tibetan Himalaya into early Early Jurassic times. They include the ammonites Choristoceras and Eopsiloceras, and the bivalves Newaagia, Terquemia, Persia, Ryderia guangdongensis, and Cultriopsis angusta. This suggests that the Tibetan Himalaya may have played a refugia role in the course of the end-Triassic mass extinction.展开更多
文摘Larger benthic foraminifera during the Eocene within the Tethyan realm played a crucial role in regional biostratigraphy.In the Middle East,particularly in Libya,Tunisia,and Egypt,nummulitids are key constituents of nummulitic limestone reservoirs,making the taxonomic identification of this structurally complex group vital for biostratigraphic and petroleum applications.This review focuses on the genus Gaziryina of the nummulitids,clarifying its taxonomic status,biostratigraphic utility,and paleobiogeographic significance using biometric data from published literature.Previously misidentified as Nummulites pulchellus,Gaziryina species have been systematically reassessed based on biometric data from published literature.The findings confirm Gaziryina as a distinct genus comprising two species:Gaziryina basatinensis(late Lutetian-Bartonian,SBZ15-18a) and Gaziryina pulchellus(late Bartonian-Priabonian,SBZ18a/b-20).Morphological characters(test shape,size progression,chamber configuration,protoconch enlargement) and ontogenetic trends(tight to lax) indicate a phylogenetic transition where Gaziryina basatinensis evolved into Gaziryina pulchellus,consistent with Cope's rule.Paleobiogeographic reconstruction indicates a probable origin in the southern NeoTethys,likely the Western Desert of Egypt,with subsequent dispersal across the northern and southeastern Tethys.These findings confirm Gaziryina as a key biostratigraphic marker,necessitating further research on its evolution and biostratigraphic framework.
基金granted by the National Natural Science Foundation of China(Grant No.41372019)the Doctoral Foundation of Ministry of Education of China(No.20110061110051)
文摘A comprehensive compilation and systematic analysis of known early and middle Permian brachiopod faunas shows that the early Permian brachiopod faunas comprise three realms,six regions,and eleven provinces,while those of the middle Permian comprise three realms,four regions,and eight provinces.A comparison and analysis of brachiopod faunal patterns reveal a coevolution between global brachiopod paleobiogeography and tectonopaleogeography during the early-middle Permian.Although temperature/latitude is the main factor controlling the formation of three realms,tectonopaleogeographic factors determine the temperature/latitude in which the continents were located.The'continental barrier'of Pangea,as a'central axis'continent,divided the three realms into six regions,which indicates that the formation of biogeographic regions was controlled mainly by the tectonopaleogeographic factors.The evolution of tectonopaleogeography was sometimes a long-term process,so that the biogeographic regions(or provinces)controlled by tectonopaleogeography displayed relative stability.Shifts in the nature of biogeographic provinces(e.g.,from cool water to warm water,and vice versa),extensions or narrowing of geographical ranges,and recombinations of some provinces were all related to regional tectonic evolution.The study of the coevolution between brachiopod paleobiogeography and tectonopaleogeography not only accounts for the formation mechanisms of brachiopod paleobiogeographic patterns during the early-middle Permian,but also provides evidences for the locations and configurations of oceans and plates(blocks)during this period.
基金supported by NSF grants ANT-1142129 to MCL, ANT-1141820 to JAC2, ANT-1142104 to PMO, ANT- 0636639 and ANT-1142052 to R. MacPhee, and OPP-9615933 and ANT-0003844 to JAC1
文摘Although the fossil record of non-avian dinosaurs from the Cretaceous of Antarctica is the poorest of any continent,fossils representing at least five major taxonomic groups(Ankylosauria,early-diverging Ornithopoda,Hadrosauridae,Titanosauria,and Theropoda)have been recovered.All come from Upper Cretaceous(Coniacian–Maastrichtian)marine and nearshore deposits belonging to the Gustav and Marambio groups of the James Ross Basin at the northern tip of the Antarctic Peninsula.The majority of these finds have come from the Campanian–Maastrichtian Snow Hill Island and López de Bertodano formations of James Ross and Vega islands.Given the rarity of Antarctic Cretaceous non-avian dinosaurs,discoveries of any fossils of these archosaurs,no matter how meager,are of significance.Here we describe fragmentary new ornithischian(ankylosaur and ornithopod)material from the upper Campanian–lower Maastrichtian Cape Lamb Member of the Snow Hill Island Formation and the Maastrichtian Sandwich Bluff Member of the López de Bertodano Formation.One of these specimens is considered to probably pertain to the holotypic individual of the early-diverging ornithopod Morrosaurus antarcticus.We also provide an up-to-date synthesis of the Late Cretaceous non-avian dinosaur record of the James Ross Basin and analyze the biostratigraphic occurrences of the various finds,demonstrating that most(including all named taxa and all reasonably complete skeletons discovered to date)occur within a relatively condensed temporal interval of the late Campanian to early Maastrichtian.Most or all James Ross Basin dinosaurs share close affinities with penecontemporaneous taxa from Patagonia,indicating that at least some continental vertebrates could disperse between southern South America and Antarctica during the final stages of the Mesozoic.
基金King Saud University for funding this work through the Researchers Supporting Project(no.RSP-2020/149)。
文摘New well-preserved rudist materials come from the Barremian–Turonian marine formations in the Yelleg, Minsherah, Maaza, and Raghawi sections in North Sinai, northern Egypt. There, 17 discovered rudist species belonging to 13 genera and seven taxonomic families are described in detail as follows: Eoradiolites plicatus(Conrad), Eoradiolites liratus(Conrad), Praeradiolites ponsianus(d'Archiac), Archaeoradiolites sp., Bournonia africana Douvillé, Bournonia fourtaui Douville, Biradiolites lombricalis(d'Orbigny), Biradiolites zumoffeni Douville, Radiolites lewyi lewyi Parnes, Radiolites sauvagesi(d'Hombres-Firmas), Durania arnaudi(Choffat), Toucasia carinata(Matheron), Toucasia sp., Neocaprina raghawiensis Steuber and Bachmann, Sellaea sp., Ichthyosarcolites sp. and Horoiopleura sp.. The Cenomanian deposits in the northern Eastern desert of Egypt only contain E. liratus. The domination of the Cenomanian rudist species in North Sinai, however, is attributed to changes in the platform, which passes mainly from carbonate in the north to siliciclastics in the south. The presence of Horiopleura sp. in the late Barremian-early Aptian deposits is documented for the first time, which suggests the expansion of taxa of the genus Horiopleura Douville to North Sinai during this interval. The age of the rudist species is documented from the late Barremian-Turonian formations, with their geographic distribution in the Mediterranean region considered..
基金Project supported by the Foundation of the Chinese National Commission for Education
文摘The distribution of ostracods and benthonic foraminifers in the China sea area is briefly reviewed from the paleobio-geographic viewpoint in this paper. Three regions can be distinguished in the area on the basis of modern distribution data: Region I (the Huanghai Sea and the Bohai Sea) with cool and temperate forms, Region I (the East China Sea and the northern part of the South China Sea) with subtropical warm-water forms and Region Ⅲ (central and southern parts of the South China Sea) with larger foraminifers and other tropical warm-water forms. The occurrence of Nummulites-Discocyclira fauna in the Eocene deposits of the East China Sea indicates a northward extension of tropical zoogeographical region at the time, whereas the distribution pattern of the Miocene Nephrolepidina-Miogypsiua-Austrotrillina fauna in the South China Sea resembles that of the present larger-foraminiferal fauna. In the South China Sea and Taiwan, a warm-water fauna with Asterorotalia and Pseudorotalia first appeared in late Miocene and then flourished in the Pliocene. Its northward invasion into the Bohai Sea area during late Pleistocene may be related to changes in current system. Many of the endemic foraminiferal and ostracod genera and species ( Sinocytheridea etc. ) in recent coastal faunas probably have originated from the South China Sea, and their occurrence in Japan or Australia may be indicative of ties between these areas during glacial times.
基金supported through TUB˙ITAK(The Scientific and Technological Research Council of Türkiye)2218—National Postdoctoral Research Fellowship Program(Project number:123C151)supported through TUB˙ITAK(2221—Fellowships for Visiting Scientists and Scientists on Sabbatical Leave)+2 种基金supported by the Beijing Natural Science Foundation(International Scientists ProjectFunding No.IS23086)by the Research Fund for International Scientists of the National Natural Science Foundation of China(Grant No.42350410443).
文摘Porcupines,members of the Hystricidae family,represent a unique group of herbivorous mammals.This study details the identification of a newly discovered mandible fragment of Hystrix primigenia,along with a right cheek tooth series from the middle Turolian Kemiklitepe-A fossil locality.While Hystrix fossils are found in numer-ous localities,the materials are often limited to a few dental fragments or isolated teeth,posing challenges to systematic investigations.The examination of this lower tooth series prompted a comprehensive review of all H.primigeniafindings across Eurasia,shedding light on its adaptive characteristics over time and space.Our paleo-biogeographical analysis indicates the absence of H.depereti in Late Miocene Türkiye,while the dispersal range of H.primigenia is broader than that of H.depereti in Eurasia.Additionally,the study delves into the discussion of H.primigenia and H.deperetifindings in Eurasia,ultimately refining the categorization of Late Miocene Hys-trix discoveries in Türkiye to two species:H.primigenia and H.kayae.Our review suggests the possibility of an additional H.kayaefinding from Samos,Greece.
文摘The Neogene interval witnessed three major waves of intercontinental dispersals by carnivoran mammals,mainly migrating from Eurasia to North America but with rare occurrences going in the opposite direction(e.g.,Sthenictis,Leptarctus).The timing and composition of dispersers are thought to be controlled by filter-bridge mechanisms,but it is unclear what types of regional conditions promoted occasional dispersals outside of the three major waves.We study and report on new occurrences of small-bodied carnivorans from late Clarendonian-aged Black Hawk Ranch and Cuyama Valley fossil mammal assemblages in California.The relatively late occurrences of the mustelids Sthenictis and Hoplictis and the ailurid Alopecocyon in coastal regions of western North America suggest that nearshore dispersal corridors both facilitate and preserve faunal elements later than they might otherwise occur further inland.The availability of both marine-and terrestrially-derived food resources may be an important factor in allowing coastally dispersing taxa to be accommodated in those ecological communities compared to less heterogenous environments further inland.
基金supported by the National Natural Science Foundation of China(4213020942002015)+3 种基金Youth Innovation Promotion Association CAS(2021070)General Project of Yunnan Provincial Science and Technology Department(202101BA070001-076)Project of Young and Middle-aged Academic and Technical Talents(202405AC350098)Meemann Chang Academician Workstation in Yunnan Province(202205AF150002)。
文摘A new species of Bothriolepis(Placodermi,Antiarcha),B.zhujiangyuanensis sp.nov.,is described from the Eifelian(Middle Devonian)Shangshuanghe Formation at the Pearl River Source Scenic Area,Qujing,Yunnan Province,southwestern China and is characterized by a broad postpineal plate bearing a straight anterior margin,a premedian plate wider than long,and a weakly developed dorsal median ridge of the trunk shield.As the eighth Bothriolepis taxon in China,B.zhujiangyuanensis sp.nov.provides a wealth of morphological data to clarify the intrageneric interrelationships.Maximum parsimony analysis,using a revised data set of the genus with 76 ingroup taxa and 72 characters,yields a well-resolved consensus tree for Bothriolepis.The phylogenetic result shows a sister-group relationship between B.zhujiangyuanensis and B.babichevi from Kazakhstan,corroborating the biogeographic proximity between the South China and Kazakhstania blocks during the Middle Devonian.
基金Financial support from the Department of Science and Technology (DST),New Delhi (Ref.no.DST/INSPIRE/03/2019/001456)supported by an INSPIRE fellowship awarded to S.K.by The Department of Science and Technology,New Delhi,INSPIRE Code (IF190496) S.K.,T.H.,M.H.
文摘Ten palm leaf impressions are documented from the latest Maastrichtian(late Cretaceous) to early Danian(earliest Paleocene) sediments(K-Pg,c.66-64 Ma) of the Mandla Lobe of the Deccan Intertrappean Beds,Madhya Pradesh,central India.The palmate leaf shape along with a definite wellpreserved costa support their placement in the subfamily Coryphoideae of the family Arecaceae.We place all recovered palm leaf specimens in the fossil genus Sabalites,report seven species of coryphoid palms and describe two new species namely,Sabalities umariaensis sp.nov.and Sabalites ghughuaensis sp.nov.The fossils indicate that coryphoid palms were highly diverse in central India by the latest Cretaceous.These and earlier reported coryphoid palm fossils from the same locality indicate that they experienced a warm and humid tropical environment during the time of deposition.These discoveries confirm the presence of a diversity of Coryphoideae in Gondwana prior to the India-Eurasia collision and provide information about coryphoid biogeographical history over geological time.Based on megafossil remains,we trace coryphoid palm migration pathways from India to mainland Southeast(SE) Asia and other parts of Asia after the docking of the Indian subcontinent with Eurasia early in the Paleogene.
文摘Ordovician conodonts representing 28 genera and 28 named and three unnamed species were identified from 740 chert and siliceous siltstone spot samples(>3000 thin sections)from deep-water turbiditic sequences of the Lachlan Orogen in central and southern New South Wales,Australia.Based on these faunas,a new conodont biozonal scheme has been established to divide the Ordovician turbiditic successions of the Lachlan Orogen into 12 superbiozones and biozones.They are(in ascending order)the Paracordylodus gracilis Superbiozone(including the Prioniodus oepiki Biozone),Periodon flabellum Superbiozone(including the Oepikodus evae Biozone in the lower part),Periodon hankensis Biozone,Periodon aculeatus Superbiozone(including the Histiodella labiosa,Histiodella holodentata,Histiodella kristinae,Pygodus serra and Pygodus anserinus biozones)and the Periodon grandis Biozone.The Pygodus anserinus Biozone is divided further into the lower and upper subbiozones.This new conodont biozonation scheme spanning the upper Tremadocian to middle Katian interval permits precise age-dating and correlation of deep-water siliciclastic rocks that characterize the Ordovician Deep-Sea Realm regionally and internationally.
文摘Recent geological and paleontological exploration in the Indus basin of Pakistan allowed the discoveries of numerous remains of non-marine reptiles( titanosaurian sauropod,abelisaurian and noasaurian theropod dinosaurs),and marine reptiles( crocodiles),flying reptiles( pterosaurs),marine and non-marine mammals,fishes,invertebrates,and plants,especially Pakistan is relatively rich in footprints / trackways in the Mesozoic.These vertebrates of Indo-Pakistan are very significant for paleobiogeographic study due to the present-day connection of this continent with Asia in Northern Hemisphere,whereas during past( Jurassic and pre-Jurassic) it was connected to the Gondwana. The Mesozoic vertebrates show close affinities with Gondwanan landmasses.The Cenozoic vertebrates show Eurasian affinity and migrated from Indo-Pak subcontinent to Eurasia or vice versa via Paleo Indus River systems along Western Indus Suture,after long journey of about 6 000 km the first collision of Indo-Pak subcontinent with Asia occurred at terminal Cretaceous.
基金supported by the Chinese Academy of Sciences (Grant Nos.XDA20070203, XDB26000000, QYZDY-SSW-DQC-22, GJHZ1885)the National Natural Science Foundation of China (Grant Nos. 41430102 and 41772018, 41625005)
文摘The first unequivocal remains of medium to large-sized mustelids from the middle Miocene Halamagai and Kekemaideng formations have been found in the Ulungur River area, Xinjiang, northwestern China. These new fossils are referred to the hypercarnivorous mustelid Hoplictis Ginsburg, 1961 and denote the first record of the genus in East Asia.We define Hoplictis baihu n. sp., for the mustelid from Tieersihabahe(Halamagai Fm.), which represents the smallest species of the genus. This primitive form is closer to H. florancei and H. noueli than to H. anatolicus and later, larger and more derived Hoplictis spp., from Europe and North America. A large toothless mandible from Duolebulejin(Kekemaiden Fm.) is assigned to Hoplictis cf. helbingi, and it presumably might represent the first record of H. helbingi outside Western Europe. The systematic position of Hoplicitis in relation to Ischyrictis with which it is similar is clarified. The occurrence in East Asia of two species of Hoplictis greatly expands the known distribution and diversity of the genus, and supports a Palaearctic Neogene dispersal event of carnivorans between Europe and Asia during the late Shanwangian–early Tunggurian equivalent to MN5–6 in Europe, and indication of another dispersal event from Europe to North America,through Northwest China during the late Tunggurian, equivalent to MN7–8 in Europe.
文摘On the basis of a worldwide review of Claraia , Pseudoclaraia and Eumorphotis , especially their important species , two range zones are recognized:the Pseudoclaraia wangi range zone —— Upper Griesbachian——and the Claraia stachei- C. aurita - Eumorphotis multiformis acme zone——Upper Griesbachian to Lower Smithian This result revises both the tradition-al Chinese concept regarding these fossils as limited in Lower Scythian and the viewpoint of some Eurasian researchers who hold Eumorphotis multiformis to be Upper Scythian .The paleogeographical distribution of the three genera is controlled primarily by temperature (latitude) and secondarily by physiographic separations .Distribution of these fossils is subdivided into four realms and five provinces . The Eastern Asia province of the Tethys realm is suggested as the source area of these fossils .The striking contrast between the composition of the species along the southern and northern margins of the Tethys , together with other distributional characters ,supports the plate tectonic theory .
文摘Numerous new records of Ferganiella, Podozamites, and Schidolepium, including a new species, Ferganiella ivantsovii sp. nov., are described from the Early Jurassic(Toarcian) Middle Subformation of the Prisayan Formation from the Euro-Sinian paleofloristic region in the Irkutsk Basin, Eastern Siberia, Russia. An analysis of the paleogeographic distribution of Ferganiella and Podozamites shows that both genera were the most diverse and numerous in the East Asian province of the Euro-Sinian region and in the Northern Chinese province of the Siberian region during the Early and Middle Jurassic. These phytochoria were located in the subtropical and temperate subtropical climate zones, which allows us to consider Ferganiella and Podozamites as thermophilic plants, which are important indicators of the Early Toarcian climatic optimum. Their abundance in the Irkutsk Basin thus may indicate Early Toarcian warming;further abundant Schidolepium cones, which produced Araucariacites pollen, typical for Euro-Sinian flora complement the scenario. Thus, the new finds are the first macrofloristic indicators of the Toarcian climatic optimum in the Irkutsk Basin.
文摘Geologic studies of the Late Cretaceous-Paleogene stratigraphic record of southern Tibet reveal distinct episodes of collision and post-collisional compression.The composition of sediments exposed along the Indus-Yarlung Zangbo suture,tectonic deformation,
文摘Saraikistan (South Punjab and surrounding) area of Pakistan is located in the central Pakistan. This area represents Triassic-Jurassic to Recent sedimentary marine and terrestrial strata. Most of the Mesozoic and Early Cenozoic are represented by marine strata with rare terrestrial deposits, while the Late Cenozoic is represented by continental fluvial deposits. This area hosts significant mineral deposits and their development can play a significant role in the development of Saraikistan region and ultimately for Pakistan. The data of recently discovered biotas from Cambrian to Miocene age are tabulated for quick view. Mesozoic biotas show a prominent paleobiogeographic link with Gondwana and Cenozoic show Eurasian. Phylogeny and hypodigm of Poripuchian titanosaurs from India and Pakistan are hinted at here.
基金financially supported by the Second Tibetan Plateau Scientific Expedition and Research Program(Grant No.2019QZKK0706)the National Natural Science Foundation of China(Grant Nos.41802002 and 91855205)the Strategic Priority Research Program(B)of Chinese Academy of Sciences(Grant No.XDB26000000)。
文摘The overall occurrence,stratigraphical distribution,palaeoenvironmental and palaeobiogeographical significances of the ostracod faunas from the Silurian Pulu Formation in the Yalai WestⅡsection,Nyalam region,southern Tibet,China,are documented for the first time.Thirty-two species belonging to 19 genera are identified and figured herein.The ostracod fauna in the Pulu Formation indicates an age of the late Llandovery-Pridoli.The ostracods belong to a podocope-rich association,which suggests an offshore environment for the Yalai West II section.The ostracods of the Pulu Formation have strong affinities with the South China taxa and also share some similarities with those from Baltica.This implies that ostracods could probably benefit from sea-level changes to facilitate faunal exchanges between peri-Gondwana and Baltica.Geographical isolation and global sea-level changes are proposed as the primary factors controlling the palaeobiogeographical distribution of ostracods during the Silurian.
基金Supported by National Natural Science Foundation of China(Grant Nos.40572013,40672012)the National Bureau of Geological Survey(Grant No.1212010818095)
文摘End-Triassic ammonoid and bivalve faunas of the Germig area, Tibetan Himalaya, lived in a tropical, shallow-water environment during the Triassic-Jurassic boundary interval. High stratigraphic resolution based on ammonite-biochrons allows to tracing the place of origin of several faunal elements. The bivalves Aguilerella and Ctenostreon occurred first in the Tibetan Himalaya and migrated from there to the eastern South Pacific, exhibiting a pantropic dispersal pattern. This dispersal route is supported by the distribution pattern of the ammonites Choristoceras, Discamphiceras, Pleuroacanthites, and Psiloceras calliphyllum. A few taxa, which went extinct everywhere else by the end of the Triassic, survived in the Tibetan Himalaya into early Early Jurassic times. They include the ammonites Choristoceras and Eopsiloceras, and the bivalves Newaagia, Terquemia, Persia, Ryderia guangdongensis, and Cultriopsis angusta. This suggests that the Tibetan Himalaya may have played a refugia role in the course of the end-Triassic mass extinction.
