Ecological stoichiometry plays an important role in revealing the mechanisms underlying biogeochemical cycles and ecosystem functions.Abiotic factors have strong effects on N-P stoichiometry,yet the impact of plant co...Ecological stoichiometry plays an important role in revealing the mechanisms underlying biogeochemical cycles and ecosystem functions.Abiotic factors have strong effects on N-P stoichiometry,yet the impact of plant community structure,especially in forests,has not been fully elucidated.We investigated 68 plots in larch forests in northern China to explore how plant community structure and environmental factors affect the N-P stoichiometry of soil and leaves.The results showed significant differences in soil and leaf N-P stoichiometry among the three larch forests,P concentration and N:P ratio of leaves were significantly related to those of soil.Except for larch forest type,N-P stoichiometry was also regulated by elevation,climatic factors,and community structure.With increasing age(from 25 to 236 years),soil N and N:P ratio significantly increased,especially in the topsoil.With increasing mean DBH,leaf N concentration and N:P ratio also increased,indicating a shift in nutrient limitations with stand growth.These findings provide evidence that plant community structure and environmental factors regulate soil and leaf N-P stoichiometry,which is critically important for understanding biogeochemical cycles and forest management undergoing natural succession.展开更多
The Dahurian larch forest in northeast China is important due to its vastness and location within a transitional zone from boreal to temperate and at the southern distribution edge of the vast Siberian larch forest. T...The Dahurian larch forest in northeast China is important due to its vastness and location within a transitional zone from boreal to temperate and at the southern distribution edge of the vast Siberian larch forest. The continuous carbon fluxes were measured from May 2004 to April 2005 in the Dahurian larch forest in Northeast China using an eddy covariance method. The results showed that the ecosystem released carbon in the dormant season from mid-October 2004 to April 2005, while it assimilated CO2 from the atmosphere in the growing season from May to September 2004. The net carbon sequestration reached its peak of 112 g.m^-2.month ^-1 in June 2004 （simplified expression of g （carbon）.m^-2.month^-1） and then gradually decreased. Annually, the larch forest was a carbon sink that sequestered carbon of 146 g-m^-2.a^-1 （simplified expression of g （carbon）.m^-2.a^-1） during the measurements. The photosynthetic process of the larch forest ecosystem was largely affected by the vapor pressure deficit （VPD） and temperature. Under humid conditions （VPD 〈 1.0 kPa）, the gross ecosystem production （GEP） increased with increasing temperature. But the net ecosystem production （NEP） showed almost no change with increasing temperature because the increment of GEP was counterbalanced by that of the ecosystem respiration. Under a dry environment （VPD 〉 1.0 kPa）, the GEP decreased with the increasing VPD at a rate of 3.0 μmol.m^-2.s^-1kPa -1 and the ecosystem respiration was also enhanced simultaneously due to the increase of air temperature, which was linearly correlated with the VPD. As a result, the net ecosystem carbon sequestration rapidly decreased with the increasing VPD at a rate of 5.2 μmol.m^-2.s-1.kPa^-1. Under humid conditions （VPD 〈 1.0 kPa）, both the GEP and NEP were obviously restricted by the low air temperature but were insensitive to the high temperature because the observed high temperature value comes within the category of the optimum range.展开更多
Background:Larch(Larix Mill.)forests are widely distributed in the upper parts of mountainous areas in China,playing vital roles in constructing mountain landscapes and maintaining mountain environments.Despite their ...Background:Larch(Larix Mill.)forests are widely distributed in the upper parts of mountainous areas in China,playing vital roles in constructing mountain landscapes and maintaining mountain environments.Despite their importance,our knowledges on the large-scale patterns of structure characteristics and the relationships between different structure variables are unclear.In this paper,we investigated 155 plots from 11 natural larch forest types across the country to explore the biogeographic patterns of the structure characteristics and the allometric relationships between different structure variables for Chinese larch forests.Results:The structure characteristics were significantly different among larch forest types.For different larch forest types,the power function fits the relationships between tree height and diameter at breast height(DBH),average DBH and stem density,and taper and stem density well,but with different exponents among larch forest types.The power exponents of the allometric relationships between tree height and DBH for different larch forest types varied from 0.61 to 0.93(mean=0.86)by standard major axis regression(SMA),and from 0.51 to 0.78(mean=0.56)by ordinary least square regression(OLS).The 50%,75%and 95%quantile regression(QR)and OLS indicated that the average DBH and taper of the L.gmelinii forests,L.gmelinii var.principis-rupprechtii forests,and L.sibirica forests were significantly correlated with stem density.Conclusions:The relationship between tree height and DBH showed a power function relationship for all larch forest types in China,but with different exponents.Overall,stem density was negatively correlated with average DBH and taper.The Sect.Larix forests exhibited stand density effect.Our findings provide an important basis for recognizing the biogeographic patterns of structure factors and for the management of larch forests in China.展开更多
Larch forests are important for species diversity,as well as soil and water conservation in mountain regions.In this study,we determined large-scale patterns of species richness in larch forests and identified the fac...Larch forests are important for species diversity,as well as soil and water conservation in mountain regions.In this study,we determined large-scale patterns of species richness in larch forests and identified the factors that drive these patterns.We found that larch forest species richness was high in southern China and low in northern China,and that patterns of species richness along an elevational gradient depend on larch forest type.In addition,we found that patterns of species richness in larch forests are best explained by contemporary climatic factors.Specifically,mean annual temperature and annual potential evapotranspiration were the most important factors for species richness of tree and shrub layers,while mean temperature of the coldest quarter and anomaly of annual precipitation from the Last Glacial Maximum to the present were the most important for that of herb layer and the whole community.Community structural factors,especially stand density,are also associated with the species richness of larch forests.Our findings that species richness in China's larch forests is mainly affected by energy availability and cold conditions support the ambient energy hypothesis and the freezing tolerance hypothesis.展开更多
Beta-diversity reflects the spatial changes in community species composition which helps to understand how communities are assembled and biodiversity is formed and maintained. Larch(Larix) forests, which are coniferou...Beta-diversity reflects the spatial changes in community species composition which helps to understand how communities are assembled and biodiversity is formed and maintained. Larch(Larix) forests, which are coniferous forests widely distributed in the mountainous and plateau areas in North and Southwest China, are critical for maintaining the environmental conditions and species diversity. Few studies of larch forests have examined the beta-diversity and its constituent components(species turnover and nestedness-resultant components). Here, we used 483 larch forest plots to determine the total betadiversity and its components in different life forms(i.e., tree, shrub, and herb) of larch forests in China and to evaluate the main drivers that underlie this beta-diversity. We found that total betadiversity of larch forests was mainly dependent on the species turnover component. In all life forms,total beta-diversity and the species turnover component increased with increasing geographic, elevational, current climatic, and paleoclimatic distances. In contrast, the nestedness-resultant component decreased across these same distances. Geographic and environmental factors explained 20%-25% of total beta-diversity, 18%-27% of species turnover component, and 4%-16% of nestedness-resultant component. Larch forest types significantly affected total beta-diversity and species turnover component. Taken together, our results indicate that life forms affect beta-diversity patterns of larch forests in China, and that beta-diversity is driven by both niche differentiation and dispersal limitation. Our findings help to greatly understand the mechanisms of community assemblies of larch forests in China.展开更多
After the analysis on the linages of species, the fractal dimensions of vegetation and soil in a managed larch(Larix gmelini) forest in Daxingan Mountains, NE China were estimated separately, and their scale correlati...After the analysis on the linages of species, the fractal dimensions of vegetation and soil in a managed larch(Larix gmelini) forest in Daxingan Mountains, NE China were estimated separately, and their scale correlation was discussed. (1) The dominant species of the larch forest, larch and the important accompany species, birch (Betula platyphylla) were homogeneously distributed along the transect. The other species were heterogeneous, and can be divided into three groups located on the right, left and central parts of the transect respectively. The transect can be separated into two parts at the position for the 29th quadrat(580m). (2) The fractal dimension in the large scale range from 200 to 400m was lower than that in the small scale range from 0 to 200m in the forest, indicating the different variation regularities of spatial heterogeneity in different scale ranges. The inflection point at 200m was one of the key scales of spatial hierarchy of the larch forest. (3) The scale variation of the forest was correlated with that of soil pH on large scales, which reflected the indirect control of the dominant species to the spatial pattern and species distribution of understorey by changing soil acidity in the forest and the indirect effects of forest management.展开更多
Thinning represents an important and frequently used silvicultural technique that improves forest wood products and has obvious effects on forest carbon stocks and stock changes. Here, we used the carbon budget model ...Thinning represents an important and frequently used silvicultural technique that improves forest wood products and has obvious effects on forest carbon stocks and stock changes. Here, we used the carbon budget model CBM-CFS3 to simulate the effects of thinning on carbon storage and changes in larch forest ecosystems under thirteen thinning scenarios. Simulation results demonstrate that strong thinning greatly reduces the biomass carbon density of larch forests compared to non-thinning forests. The minimum and maximum average biomass carbon density, respectively, were 30.3 tC ha^-1 and 47.8 tC ha^-1, a difference of 58% under set scenarios in the simulated time scale. The dead organic matter（including soil） carbon density increased in all thinned larch forests stands, compared with non-thinning stands, and the pattern of variation was opposite to that found for biomass carbon density. However, the total ecosystem carbon density of larch forests declined with thinning because the increase in dead organic matter carbon is insufficient to offset the loss of biomass carbon caused by thinning. Thus, strong thinning can transform larch forest ecosystems from carbon sinks into carbon sources. Future work should consider the carbon sequestered in wood materials acquired via thinning and their use as substitutes for other construction materials with less favorable lifecycle carbon footprints, such as steel, cement, aluminum and PVC.展开更多
Soils were collected from.three neighboring forest sites: 36-year-old larch plantation, 11-year-old larch plantation, and natural secondary broad-leaved forest (as control). Soil pH, total C. totaI N, C/N ratio. and ...Soils were collected from.three neighboring forest sites: 36-year-old larch plantation, 11-year-old larch plantation, and natural secondary broad-leaved forest (as control). Soil pH, total C. totaI N, C/N ratio. and available N (NO3-N and NH4-N) were measured. Laboratory incubations of soil samples were conducted during a 50 days period for the measurement of nitrogen mineralization rate and nitrification potenial. The results proved a degeneration in soil nitrogen status with stand age of larch plantations, which implicated one important aspect of soil degradation when natural forest was replaced by coniferous plantations.展开更多
Allelopathic effect of larch (Larix gmelini ) on the ash growth (Fraximus mandshurica) was studied in artificial cultivation tests. The results revealed that the larch root secretion obviously stimulated the ash growt...Allelopathic effect of larch (Larix gmelini ) on the ash growth (Fraximus mandshurica) was studied in artificial cultivation tests. The results revealed that the larch root secretion obviously stimulated the ash growth. In order to determine the main stimulation allelochemicals, the chemical composition was analyzed. By contrasting the contents of carbohydrate and aminoacid in root secretion of larch and ash, it was concluded that the carbohydrate and aminoacid were not important stimulation allelochemicals. The organic acid and other components in root secration of larch and ash were analyzed by GC and GC-MS analysis. The sand culture tests were carried out ed selected model compounds. The results showed that benzeneacetic acid, benzenepropionic acid and phenolic acids in root secretion of larch were the main stimulation allelochemicals.展开更多
According to the study on soils under the 34-year-old Larch forest and Larch-Walnut mixed forest. It tvas concluded that, in mixed forest, the total content of water- stable aggregate (0. 25- 5mm) in the upper layers ...According to the study on soils under the 34-year-old Larch forest and Larch-Walnut mixed forest. It tvas concluded that, in mixed forest, the total content of water- stable aggregate (0. 25- 5mm) in the upper layers (0- 30cm) of soil was significant ly higher than that in pure forest. The 2mm aggregates increased 48% in mixed forest soil compared with pure fOrest. and consequently, the three-phase ratio of soil was regulate d, the physical properties improved and the fertility of soil raised. Because of the stability of total aggregate content and the importance of 2mm aggregate content in soil fertility.it s advisable using the two indices above to characterize the effect of mixed forest on soil improvement.展开更多
Allelopathic effect of larch (Larix gmelini) leaching on ash (Fraxinus mandshurica) growth was studied with sand culture test. It was understood that the ash growth was stimulated by the larch leaching. The chemical c...Allelopathic effect of larch (Larix gmelini) leaching on ash (Fraxinus mandshurica) growth was studied with sand culture test. It was understood that the ash growth was stimulated by the larch leaching. The chemical compositions of the larch and ash leaching were identified and determined by chemical and instrumental analysis. The results showed that in the larch and ash leaching there were fatty acids (including monocarboxylic, dicarboxylic and tricarboxylic acids), phenolic acids, tannin, aromatic alcohols, polylol, aminoacids and carbohydrate, etc. on the basis of the analytical results of chemical composition, with reference to the results of model compound test, it is considered that the citric acid in the larch leaching was the main simulative allelochemical to the ash growth, and trihydroxybenzeneacetic acid was the subordinate stimulative allelochemical.展开更多
With the method of stem analyses by the microscope, the xylem’s accumulations atdifferent ages were quantitatively studied using the data of typical plant association for Dahurianlarch (Larix gmelini Rupr) and Manchu...With the method of stem analyses by the microscope, the xylem’s accumulations atdifferent ages were quantitatively studied using the data of typical plant association for Dahurianlarch (Larix gmelini Rupr) and Manchurian ash (Fraxinus mandshurica Rupr) mixed forests. Thedeveloping process for this important mixed forest type in the study area was systematically established. The result showed that in the artificial stand of Dahurian larch mixed with naturalManchurian ash. only those which have the same origin as Dahurian larch can grow up before theclosing of stand and form the components of the main layers. After 24 years seif-thinning, when theywere 29 years old, Dahurian larch would be at the stage of no self-thinning. At last the stand woulddevelop to the broad-leaved forests dominating with the Manchurian ash.展开更多
For evaluating microbial community changes in a Siberian larch stand disturbed by forest fire or clearcutting,357 clones were randomly selected and sequenced using a culture-independent approach and 16S rRNA sequencin...For evaluating microbial community changes in a Siberian larch stand disturbed by forest fire or clearcutting,357 clones were randomly selected and sequenced using a culture-independent approach and 16S rRNA sequencing to characterize the bacterial composition and diversity from the different disturbed Siberian larch stands.Interestingly,the burned larch stand had an increase in the relative amounts of b-proteobacteria and Firmicutes and a decrease in Acidobacteria,while Gemmatimonadetes increased Verrucomicrobia decreased in the harvested larch stand.Microbial diversity and richness were higher in the undisturbed larch stand than the disturbed(burned or clear-cut)larch stands,and the influence of clear-cutting was more negative than that of the forest fire.This study indicates that evaluating the microbial diversity of undisturbed,burned,and clear-cutting Siberian larch stands provides information about the impact of forest disturbances on soil microbial communities,which may be helpful for understanding and evaluating soil health and devising reafforestation strategies for larch.展开更多
Due to increasing timber demands, large areas of secondary forests have been converted to larch plantations （LPs） in Northeast China because the secondary forests could not produce timber as much as LPs. However, th...Due to increasing timber demands, large areas of secondary forests have been converted to larch plantations （LPs） in Northeast China because the secondary forests could not produce timber as much as LPs. However, there are a series of ecological problems such as lower soil fertility, reduced water-holding capacity and acidification of surface runoff water occurring in LPs because of the single-species composition of LPs. Therefore, a guidance on how to transform LPs into larch-broadleaf mixed forests （LBMFs） at a large spatial scale is needed for local foresters. First, Landsat time series data set and SPOT-5 images were combined to map the spatial-temporal distribution of LPs in Northeast China. Then, the topographical characteristics of LPs in 2010s were determined. Furthermore, three sub-regions of LPs were divided closely linking to their ecosystem services and forest management aims. Finally, detailed information on how to transform the LPs into LBMFs was given according to the three sub-regions. The results showed that the area of LPs increased during 1980s and 2010s, and reached 2.61 million ha in 2010s. Of which, 0.72 million ha （27.6%） and 1.89 million ha （72.4%） LPs distributed in slopes less than 5° and greater than 5°, respectively. Of the LPs （72.4%） in slopes greater than 5°, 48.7 and 23.7% located in downslope （LPs locating at the down slope of adjacent secondary forests in the same aspect） and upslope （LPs locating at the up slope of adjacent secondary forests in the same aspect）, respectively; 0.10 million ha （3.8%） located in slopes greater than 25°. The LPs were divided into Sub-Regions-I, II, III according to slopes. For Sub-region-I （top role is to produce timber）, self-fertilizing shrub species can be intro- duced after clean cutting for young-aged LPs, and release thinning can be applied for middle-aged LPs, fast growth thinning for near-matured LPs, and clear cutting for matured LPs. For Sub-Region-II （the priority is to provide water conservation combined with timber production）, the LPs should be induced into LBMFs. The LPs located the downslope positions in Sub-Region-II can be transformed to LBMFs by natural regeneration of broadleaved tree species after thinning because of enough seed sources of broadleaved tree species from the secondary forests locating the upslope positions. The LPs located the upslope positions or the region where seed sources of broadleaved tree species are unavailable in Sub-Region-II must be induced to LBMFs by al＇tificial regeneration. For Sub-Re- gion-III （the preference is only for water conservation because the slope is greater than 25°）, the LPs should be particularly protected from intensive disturbances, and induced into LBMFs by natural regeneration.展开更多
For the demands of automatic pruning, an effective discrimination rule of the forked and non-forked larch trees is established. First, information of tnmk and branch diameters of a larch plantations was collected from...For the demands of automatic pruning, an effective discrimination rule of the forked and non-forked larch trees is established. First, information of tnmk and branch diameters of a larch plantations was collected from the west mountain of Beijing. The growth characteristics of the forked and non-forked trees were studied. Given the statistical characteristics of the trunk and branch diameters, a discriminant function of the forked branch and non-forked larch trees was established statistically. Excellent discrimination results were obtained by the function and the rule. The study presents an effective discrimination rule to separate forked trees from straight trees for automatic pruning.展开更多
Biomass functions were established to estimate above-ground biomass of Siberian larch （Larix sibirica） in the Altai Mountains of Mon- golia. The functions are based on biomass sampling of trees from 18 different sit...Biomass functions were established to estimate above-ground biomass of Siberian larch （Larix sibirica） in the Altai Mountains of Mon- golia. The functions are based on biomass sampling of trees from 18 different sites, which represent the driest locations within the natural range ofL. sibirica. The best performing regression model was found for the equations y = （D2 H）/（a＋bD） for stem biomass, y = aDb for branch biomass, and y=aDb Hc for needle biomass, where D is the stem diameter at breast height and H is the tree height. The robustness of the biomass functions is assessed by comparison with equations which had been previously published from a plantation in Iceland. There, y=aDb Hc was found to be the most significant model for stem and total above-ground biomasses. Applying the equations from Iceland for estimating the above-ground biomass of trees from Mongolia resulted in the underesti- mation of the biomass in large-diameter trees and the overestimation of the biomass in thin trees. The underestimation of thick-stemmed trees is probably attributable to the higher wood density, which has to be ex- pected under the ultracontinental climate of Mongolia compared to the euoceanic climate of Iceland. The overestimation of the biomass in trees with low stem diameter is probably due to the high density of young growth in the not systematically managed forests of the Mongolian Altai Mountains, which inhibits branching, whereas the plantations in Iceland are likely to have been planted in lower densities.展开更多
Carbon （C） dynamics are central to understanding ecosystem restoration effects within the context of Grain for Green Project （GGP）. GGP stared in China since 2003 to improve the environment. Despite its importance...Carbon （C） dynamics are central to understanding ecosystem restoration effects within the context of Grain for Green Project （GGP）. GGP stared in China since 2003 to improve the environment. Despite its importance, how total forest ecosystem C stock （FECS） develops fol- lowing land-use changes from cropland to plantation is poorly under- stood, in particular the relationship of C allocation to pools. We quanti- fied C pools in a chronosequence ranging from 0 to 48 years, using com- plete above- and below-ground harvests based on detailed field inventory Stands were chosen along a succession sequence in managed plantations of Korean larch （Larix olgensis Henry.）, a native planting species in the Lesser Khingan Mountains, Northeast of China. The FECS of Korean larch plantation （KLP） were dynamic across stand development, chang- ing from 88.2 Mg.ha-1 at cropland, to 183.9 Mg.ha·-1 as an average of forest C from 7- through 48-year-old plantation. In a 48-year-old mature KLP, vegetation comprises 48.63% of FECS and accounts for 67.66% of annual net C increment （ANCI）. Soil is responsible for 38.19% and 13.53% of those, and with the remainders of 13.18% and 18.81% in down woody materials. Based on comparisons of our estimate to those of others, we conclude that afforestation of Korean larch plantation is a valid approach to sequester carbon.展开更多
Permanent sample plots were set up for researching interspecific relationship of the artificia1 larch stand mixed with natural ash in the slash site in Maoershan Experiment Forestry Farm of Northeast Forestry Universi...Permanent sample plots were set up for researching interspecific relationship of the artificia1 larch stand mixed with natural ash in the slash site in Maoershan Experiment Forestry Farm of Northeast Forestry University. All trees in the plots were taken as objective trees and tree number,DBH, distance between trees and that of crown diameter were measured in range of 1-6m from every objective tree. Then study by Alemdag competition index, variance analysis and quantitation method were made. The result showed that the intraspecific competition was more intensive than the interspecific competition. Especially the competition between larch and ash was least relatively. For these kinds of site, the suitable mixture at 20-30 ages should be 2800 Ind/hm2 in density and 2 larch;1 ash in mixed ratio, which might improve site condition to promote stand produetivity.展开更多
基金supported by the National Natural Science Foundation of China(No.32201426,No.31988102)the Major Program for Basic Research Project of Yunnan Province(No.202101BC070002)the Key Research and Development Program of Yunnan Provin ce(No.202303AC100009).
文摘Ecological stoichiometry plays an important role in revealing the mechanisms underlying biogeochemical cycles and ecosystem functions.Abiotic factors have strong effects on N-P stoichiometry,yet the impact of plant community structure,especially in forests,has not been fully elucidated.We investigated 68 plots in larch forests in northern China to explore how plant community structure and environmental factors affect the N-P stoichiometry of soil and leaves.The results showed significant differences in soil and leaf N-P stoichiometry among the three larch forests,P concentration and N:P ratio of leaves were significantly related to those of soil.Except for larch forest type,N-P stoichiometry was also regulated by elevation,climatic factors,and community structure.With increasing age(from 25 to 236 years),soil N and N:P ratio significantly increased,especially in the topsoil.With increasing mean DBH,leaf N concentration and N:P ratio also increased,indicating a shift in nutrient limitations with stand growth.These findings provide evidence that plant community structure and environmental factors regulate soil and leaf N-P stoichiometry,which is critically important for understanding biogeochemical cycles and forest management undergoing natural succession.
基金the Global Environment Research Fund,Ministry of the Environment,Japan (S-1: Integrated Study for Terrestrial Carbon Management of Asia in the 21st Century Based on Scientific Advancements)the Chinese Academy of Sciences (07W70000SZ)+1 种基金the National Natural Science Foundation of China (30300271)the State Key Basic Research and Development Plan of China (2004CCA02700)
文摘The Dahurian larch forest in northeast China is important due to its vastness and location within a transitional zone from boreal to temperate and at the southern distribution edge of the vast Siberian larch forest. The continuous carbon fluxes were measured from May 2004 to April 2005 in the Dahurian larch forest in Northeast China using an eddy covariance method. The results showed that the ecosystem released carbon in the dormant season from mid-October 2004 to April 2005, while it assimilated CO2 from the atmosphere in the growing season from May to September 2004. The net carbon sequestration reached its peak of 112 g.m^-2.month ^-1 in June 2004 （simplified expression of g （carbon）.m^-2.month^-1） and then gradually decreased. Annually, the larch forest was a carbon sink that sequestered carbon of 146 g-m^-2.a^-1 （simplified expression of g （carbon）.m^-2.a^-1） during the measurements. The photosynthetic process of the larch forest ecosystem was largely affected by the vapor pressure deficit （VPD） and temperature. Under humid conditions （VPD 〈 1.0 kPa）, the gross ecosystem production （GEP） increased with increasing temperature. But the net ecosystem production （NEP） showed almost no change with increasing temperature because the increment of GEP was counterbalanced by that of the ecosystem respiration. Under a dry environment （VPD 〉 1.0 kPa）, the GEP decreased with the increasing VPD at a rate of 3.0 μmol.m^-2.s^-1kPa -1 and the ecosystem respiration was also enhanced simultaneously due to the increase of air temperature, which was linearly correlated with the VPD. As a result, the net ecosystem carbon sequestration rapidly decreased with the increasing VPD at a rate of 5.2 μmol.m^-2.s-1.kPa^-1. Under humid conditions （VPD 〈 1.0 kPa）, both the GEP and NEP were obviously restricted by the low air temperature but were insensitive to the high temperature because the observed high temperature value comes within the category of the optimum range.
基金the National Science and Technology Basic Project of China(No.2015FY210200)National Natural Science Foundation(No.31988102).
文摘Background:Larch(Larix Mill.)forests are widely distributed in the upper parts of mountainous areas in China,playing vital roles in constructing mountain landscapes and maintaining mountain environments.Despite their importance,our knowledges on the large-scale patterns of structure characteristics and the relationships between different structure variables are unclear.In this paper,we investigated 155 plots from 11 natural larch forest types across the country to explore the biogeographic patterns of the structure characteristics and the allometric relationships between different structure variables for Chinese larch forests.Results:The structure characteristics were significantly different among larch forest types.For different larch forest types,the power function fits the relationships between tree height and diameter at breast height(DBH),average DBH and stem density,and taper and stem density well,but with different exponents among larch forest types.The power exponents of the allometric relationships between tree height and DBH for different larch forest types varied from 0.61 to 0.93(mean=0.86)by standard major axis regression(SMA),and from 0.51 to 0.78(mean=0.56)by ordinary least square regression(OLS).The 50%,75%and 95%quantile regression(QR)and OLS indicated that the average DBH and taper of the L.gmelinii forests,L.gmelinii var.principis-rupprechtii forests,and L.sibirica forests were significantly correlated with stem density.Conclusions:The relationship between tree height and DBH showed a power function relationship for all larch forest types in China,but with different exponents.Overall,stem density was negatively correlated with average DBH and taper.The Sect.Larix forests exhibited stand density effect.Our findings provide an important basis for recognizing the biogeographic patterns of structure factors and for the management of larch forests in China.
基金supported by the National Natural Science Foundation of China(No.31988102)the National Science and Technology Basic Project of China(No.2015FY210200).
文摘Larch forests are important for species diversity,as well as soil and water conservation in mountain regions.In this study,we determined large-scale patterns of species richness in larch forests and identified the factors that drive these patterns.We found that larch forest species richness was high in southern China and low in northern China,and that patterns of species richness along an elevational gradient depend on larch forest type.In addition,we found that patterns of species richness in larch forests are best explained by contemporary climatic factors.Specifically,mean annual temperature and annual potential evapotranspiration were the most important factors for species richness of tree and shrub layers,while mean temperature of the coldest quarter and anomaly of annual precipitation from the Last Glacial Maximum to the present were the most important for that of herb layer and the whole community.Community structural factors,especially stand density,are also associated with the species richness of larch forests.Our findings that species richness in China's larch forests is mainly affected by energy availability and cold conditions support the ambient energy hypothesis and the freezing tolerance hypothesis.
基金supported by the Major Program for Basic Research Project of Yunnan Province (No. 202101BC070002)the National Natural Science Foundation of China (No. 32201426, No. 31988102)the National Science and Technology Basic Project of China (No. 2015FY210200)
文摘Beta-diversity reflects the spatial changes in community species composition which helps to understand how communities are assembled and biodiversity is formed and maintained. Larch(Larix) forests, which are coniferous forests widely distributed in the mountainous and plateau areas in North and Southwest China, are critical for maintaining the environmental conditions and species diversity. Few studies of larch forests have examined the beta-diversity and its constituent components(species turnover and nestedness-resultant components). Here, we used 483 larch forest plots to determine the total betadiversity and its components in different life forms(i.e., tree, shrub, and herb) of larch forests in China and to evaluate the main drivers that underlie this beta-diversity. We found that total betadiversity of larch forests was mainly dependent on the species turnover component. In all life forms,total beta-diversity and the species turnover component increased with increasing geographic, elevational, current climatic, and paleoclimatic distances. In contrast, the nestedness-resultant component decreased across these same distances. Geographic and environmental factors explained 20%-25% of total beta-diversity, 18%-27% of species turnover component, and 4%-16% of nestedness-resultant component. Larch forest types significantly affected total beta-diversity and species turnover component. Taken together, our results indicate that life forms affect beta-diversity patterns of larch forests in China, and that beta-diversity is driven by both niche differentiation and dispersal limitation. Our findings help to greatly understand the mechanisms of community assemblies of larch forests in China.
基金TheNationalNaturalScienceFoundationofChina (No .39770 1 50and 39470 1 2 9)
文摘After the analysis on the linages of species, the fractal dimensions of vegetation and soil in a managed larch(Larix gmelini) forest in Daxingan Mountains, NE China were estimated separately, and their scale correlation was discussed. (1) The dominant species of the larch forest, larch and the important accompany species, birch (Betula platyphylla) were homogeneously distributed along the transect. The other species were heterogeneous, and can be divided into three groups located on the right, left and central parts of the transect respectively. The transect can be separated into two parts at the position for the 29th quadrat(580m). (2) The fractal dimension in the large scale range from 200 to 400m was lower than that in the small scale range from 0 to 200m in the forest, indicating the different variation regularities of spatial heterogeneity in different scale ranges. The inflection point at 200m was one of the key scales of spatial hierarchy of the larch forest. (3) The scale variation of the forest was correlated with that of soil pH on large scales, which reflected the indirect control of the dominant species to the spatial pattern and species distribution of understorey by changing soil acidity in the forest and the indirect effects of forest management.
基金Strategic Priority Research Program of Chinese Academy of Sciences(XDA05050203)Knowledge Innovation Projects of Significant Directions of Chinese Academy of Sciences(KZCX2-YW-Q1-05)
文摘Thinning represents an important and frequently used silvicultural technique that improves forest wood products and has obvious effects on forest carbon stocks and stock changes. Here, we used the carbon budget model CBM-CFS3 to simulate the effects of thinning on carbon storage and changes in larch forest ecosystems under thirteen thinning scenarios. Simulation results demonstrate that strong thinning greatly reduces the biomass carbon density of larch forests compared to non-thinning forests. The minimum and maximum average biomass carbon density, respectively, were 30.3 tC ha^-1 and 47.8 tC ha^-1, a difference of 58% under set scenarios in the simulated time scale. The dead organic matter（including soil） carbon density increased in all thinned larch forests stands, compared with non-thinning stands, and the pattern of variation was opposite to that found for biomass carbon density. However, the total ecosystem carbon density of larch forests declined with thinning because the increase in dead organic matter carbon is insufficient to offset the loss of biomass carbon caused by thinning. Thus, strong thinning can transform larch forest ecosystems from carbon sinks into carbon sources. Future work should consider the carbon sequestered in wood materials acquired via thinning and their use as substitutes for other construction materials with less favorable lifecycle carbon footprints, such as steel, cement, aluminum and PVC.
文摘Soils were collected from.three neighboring forest sites: 36-year-old larch plantation, 11-year-old larch plantation, and natural secondary broad-leaved forest (as control). Soil pH, total C. totaI N, C/N ratio. and available N (NO3-N and NH4-N) were measured. Laboratory incubations of soil samples were conducted during a 50 days period for the measurement of nitrogen mineralization rate and nitrification potenial. The results proved a degeneration in soil nitrogen status with stand age of larch plantations, which implicated one important aspect of soil degradation when natural forest was replaced by coniferous plantations.
文摘Allelopathic effect of larch (Larix gmelini ) on the ash growth (Fraximus mandshurica) was studied in artificial cultivation tests. The results revealed that the larch root secretion obviously stimulated the ash growth. In order to determine the main stimulation allelochemicals, the chemical composition was analyzed. By contrasting the contents of carbohydrate and aminoacid in root secretion of larch and ash, it was concluded that the carbohydrate and aminoacid were not important stimulation allelochemicals. The organic acid and other components in root secration of larch and ash were analyzed by GC and GC-MS analysis. The sand culture tests were carried out ed selected model compounds. The results showed that benzeneacetic acid, benzenepropionic acid and phenolic acids in root secretion of larch were the main stimulation allelochemicals.
文摘According to the study on soils under the 34-year-old Larch forest and Larch-Walnut mixed forest. It tvas concluded that, in mixed forest, the total content of water- stable aggregate (0. 25- 5mm) in the upper layers (0- 30cm) of soil was significant ly higher than that in pure forest. The 2mm aggregates increased 48% in mixed forest soil compared with pure fOrest. and consequently, the three-phase ratio of soil was regulate d, the physical properties improved and the fertility of soil raised. Because of the stability of total aggregate content and the importance of 2mm aggregate content in soil fertility.it s advisable using the two indices above to characterize the effect of mixed forest on soil improvement.
文摘Allelopathic effect of larch (Larix gmelini) leaching on ash (Fraxinus mandshurica) growth was studied with sand culture test. It was understood that the ash growth was stimulated by the larch leaching. The chemical compositions of the larch and ash leaching were identified and determined by chemical and instrumental analysis. The results showed that in the larch and ash leaching there were fatty acids (including monocarboxylic, dicarboxylic and tricarboxylic acids), phenolic acids, tannin, aromatic alcohols, polylol, aminoacids and carbohydrate, etc. on the basis of the analytical results of chemical composition, with reference to the results of model compound test, it is considered that the citric acid in the larch leaching was the main simulative allelochemical to the ash growth, and trihydroxybenzeneacetic acid was the subordinate stimulative allelochemical.
文摘With the method of stem analyses by the microscope, the xylem’s accumulations atdifferent ages were quantitatively studied using the data of typical plant association for Dahurianlarch (Larix gmelini Rupr) and Manchurian ash (Fraxinus mandshurica Rupr) mixed forests. Thedeveloping process for this important mixed forest type in the study area was systematically established. The result showed that in the artificial stand of Dahurian larch mixed with naturalManchurian ash. only those which have the same origin as Dahurian larch can grow up before theclosing of stand and form the components of the main layers. After 24 years seif-thinning, when theywere 29 years old, Dahurian larch would be at the stage of no self-thinning. At last the stand woulddevelop to the broad-leaved forests dominating with the Manchurian ash.
基金supported by Daegu University Research Grant(No.20130508)
文摘For evaluating microbial community changes in a Siberian larch stand disturbed by forest fire or clearcutting,357 clones were randomly selected and sequenced using a culture-independent approach and 16S rRNA sequencing to characterize the bacterial composition and diversity from the different disturbed Siberian larch stands.Interestingly,the burned larch stand had an increase in the relative amounts of b-proteobacteria and Firmicutes and a decrease in Acidobacteria,while Gemmatimonadetes increased Verrucomicrobia decreased in the harvested larch stand.Microbial diversity and richness were higher in the undisturbed larch stand than the disturbed(burned or clear-cut)larch stands,and the influence of clear-cutting was more negative than that of the forest fire.This study indicates that evaluating the microbial diversity of undisturbed,burned,and clear-cutting Siberian larch stands provides information about the impact of forest disturbances on soil microbial communities,which may be helpful for understanding and evaluating soil health and devising reafforestation strategies for larch.
基金supported by the National Basic Research Program of China(973 Program,No.2012CB416906)
文摘Due to increasing timber demands, large areas of secondary forests have been converted to larch plantations （LPs） in Northeast China because the secondary forests could not produce timber as much as LPs. However, there are a series of ecological problems such as lower soil fertility, reduced water-holding capacity and acidification of surface runoff water occurring in LPs because of the single-species composition of LPs. Therefore, a guidance on how to transform LPs into larch-broadleaf mixed forests （LBMFs） at a large spatial scale is needed for local foresters. First, Landsat time series data set and SPOT-5 images were combined to map the spatial-temporal distribution of LPs in Northeast China. Then, the topographical characteristics of LPs in 2010s were determined. Furthermore, three sub-regions of LPs were divided closely linking to their ecosystem services and forest management aims. Finally, detailed information on how to transform the LPs into LBMFs was given according to the three sub-regions. The results showed that the area of LPs increased during 1980s and 2010s, and reached 2.61 million ha in 2010s. Of which, 0.72 million ha （27.6%） and 1.89 million ha （72.4%） LPs distributed in slopes less than 5° and greater than 5°, respectively. Of the LPs （72.4%） in slopes greater than 5°, 48.7 and 23.7% located in downslope （LPs locating at the down slope of adjacent secondary forests in the same aspect） and upslope （LPs locating at the up slope of adjacent secondary forests in the same aspect）, respectively; 0.10 million ha （3.8%） located in slopes greater than 25°. The LPs were divided into Sub-Regions-I, II, III according to slopes. For Sub-region-I （top role is to produce timber）, self-fertilizing shrub species can be intro- duced after clean cutting for young-aged LPs, and release thinning can be applied for middle-aged LPs, fast growth thinning for near-matured LPs, and clear cutting for matured LPs. For Sub-Region-II （the priority is to provide water conservation combined with timber production）, the LPs should be induced into LBMFs. The LPs located the downslope positions in Sub-Region-II can be transformed to LBMFs by natural regeneration of broadleaved tree species after thinning because of enough seed sources of broadleaved tree species from the secondary forests locating the upslope positions. The LPs located the upslope positions or the region where seed sources of broadleaved tree species are unavailable in Sub-Region-II must be induced to LBMFs by al＇tificial regeneration. For Sub-Re- gion-III （the preference is only for water conservation because the slope is greater than 25°）, the LPs should be particularly protected from intensive disturbances, and induced into LBMFs by natural regeneration.
文摘For the demands of automatic pruning, an effective discrimination rule of the forked and non-forked larch trees is established. First, information of tnmk and branch diameters of a larch plantations was collected from the west mountain of Beijing. The growth characteristics of the forked and non-forked trees were studied. Given the statistical characteristics of the trunk and branch diameters, a discriminant function of the forked branch and non-forked larch trees was established statistically. Excellent discrimination results were obtained by the function and the rule. The study presents an effective discrimination rule to separate forked trees from straight trees for automatic pruning.
基金funded by the Asian Research Center (ARC) based in the National University of Mongolia (Ulan Bator) The work was done in association with the project "Forest regeneration and biodiversity at the forest-steppe border of the Altai and Khangai Mountains under contrasting developments of livestock numbers in Kazakhstan and Mongolia" funded by the Volkswagen Foundation
文摘Biomass functions were established to estimate above-ground biomass of Siberian larch （Larix sibirica） in the Altai Mountains of Mon- golia. The functions are based on biomass sampling of trees from 18 different sites, which represent the driest locations within the natural range ofL. sibirica. The best performing regression model was found for the equations y = （D2 H）/（a＋bD） for stem biomass, y = aDb for branch biomass, and y=aDb Hc for needle biomass, where D is the stem diameter at breast height and H is the tree height. The robustness of the biomass functions is assessed by comparison with equations which had been previously published from a plantation in Iceland. There, y=aDb Hc was found to be the most significant model for stem and total above-ground biomasses. Applying the equations from Iceland for estimating the above-ground biomass of trees from Mongolia resulted in the underesti- mation of the biomass in large-diameter trees and the overestimation of the biomass in thin trees. The underestimation of thick-stemmed trees is probably attributable to the higher wood density, which has to be ex- pected under the ultracontinental climate of Mongolia compared to the euoceanic climate of Iceland. The overestimation of the biomass in trees with low stem diameter is probably due to the high density of young growth in the not systematically managed forests of the Mongolian Altai Mountains, which inhibits branching, whereas the plantations in Iceland are likely to have been planted in lower densities.
基金supported by the Special Public Interest Research and Industry Fund of Forestry(No.200904003-1)Project of Forestry Science and Technology Research(No.2012-07)the Importation of Foreign Advanced Agricultural Science and Technology Program(2008-4-48)
文摘Carbon （C） dynamics are central to understanding ecosystem restoration effects within the context of Grain for Green Project （GGP）. GGP stared in China since 2003 to improve the environment. Despite its importance, how total forest ecosystem C stock （FECS） develops fol- lowing land-use changes from cropland to plantation is poorly under- stood, in particular the relationship of C allocation to pools. We quanti- fied C pools in a chronosequence ranging from 0 to 48 years, using com- plete above- and below-ground harvests based on detailed field inventory Stands were chosen along a succession sequence in managed plantations of Korean larch （Larix olgensis Henry.）, a native planting species in the Lesser Khingan Mountains, Northeast of China. The FECS of Korean larch plantation （KLP） were dynamic across stand development, chang- ing from 88.2 Mg.ha-1 at cropland, to 183.9 Mg.ha·-1 as an average of forest C from 7- through 48-year-old plantation. In a 48-year-old mature KLP, vegetation comprises 48.63% of FECS and accounts for 67.66% of annual net C increment （ANCI）. Soil is responsible for 38.19% and 13.53% of those, and with the remainders of 13.18% and 18.81% in down woody materials. Based on comparisons of our estimate to those of others, we conclude that afforestation of Korean larch plantation is a valid approach to sequester carbon.
文摘Permanent sample plots were set up for researching interspecific relationship of the artificia1 larch stand mixed with natural ash in the slash site in Maoershan Experiment Forestry Farm of Northeast Forestry University. All trees in the plots were taken as objective trees and tree number,DBH, distance between trees and that of crown diameter were measured in range of 1-6m from every objective tree. Then study by Alemdag competition index, variance analysis and quantitation method were made. The result showed that the intraspecific competition was more intensive than the interspecific competition. Especially the competition between larch and ash was least relatively. For these kinds of site, the suitable mixture at 20-30 ages should be 2800 Ind/hm2 in density and 2 larch;1 ash in mixed ratio, which might improve site condition to promote stand produetivity.
