A set of standard chronologies for tree-ring width(TRW),earlywood width(EWW)and latewood width(LWW)in Pinus tabuliformis Carr.along an altitudi-nal gradient(1450,1400,and 1350 m a.s.l.)on Baiyunshan Mountain,Central C...A set of standard chronologies for tree-ring width(TRW),earlywood width(EWW)and latewood width(LWW)in Pinus tabuliformis Carr.along an altitudi-nal gradient(1450,1400,and 1350 m a.s.l.)on Baiyunshan Mountain,Central China to analyze the effect of varying temperature and precipitation on growth along the gradi-ent.Correlation analyses showed that at all three altitudes and the TRW and EWW chronologies generally had signifi-cant negative correlations with mean and maximum tem-peratures in the current April and May and with minimum temperatures in the prior July and August,but significant positive correlations with precipitation in the current May.Correlations were generally significantly negative between LWW chronologies and all temperatures in the prior July and August,indicating that the prior summer temperature had a strong lag effect on the growth of P.tabuliformis that increased with altitude.The correlation with the standard-ized precipitation evapotranspiration index(SPEI)confirmed that wet conditions in the current May promoted growth of TR and EW at all altitudes.Significant altitudinal differences were also found;at 1400 m,there were significant positive correlations between EWW chronologies and SPEI in the current April and significant negative correlations between LWW chronologies and SPEI in the current September,but these correlations were not significant at 1450 m.At 1350 m,there were also significant negative correlations between the TRW and the EWW chronologies and SPEI in the prior October and the current July and between LWW chronology and SPEI in the current August,but these cor-relations were not significant at 1400 m.Moving correlation results showed a stable response of EWW in relation to the SPEI in the current May at all three altitudes and of LWW to maximum temperature in the prior July-August at 1400 m from 2002 to 2018.The EWW chronology at 1400 m and the LWW chronology at 1450 m were identified as more suitable for climate reconstruction.These results provide a strong scientific basis for forest management decisions and climate reconstructions in Central China.展开更多
Temporal changes in the relationship between tree growth and climate have been observed in numerous forests across the world.The patterns and the possible regu-lators(e.g.,forest community structure)of such changes ar...Temporal changes in the relationship between tree growth and climate have been observed in numerous forests across the world.The patterns and the possible regu-lators(e.g.,forest community structure)of such changes are,however,not well understood.A vegetation survey and analyses of growth-climate relationships for Abies georgei var.Smithii(Smith fir)forests were carried along an altitudi-nal gradient from 3600 to 4200 m on Meili Snow Mountain,southeastern Tibetan Plateau.The results showed that the associations between growth and temperature have declined since the 1970s over the whole transect,while response to standardized precipitation-evapotranspiration indices(SPEI)strengthened in the mid-and lower-transect.Comparison between growth and vegetation data showed that tree growth was more sensitive to drought in stands with higher species richness and greater shrub cover.Drought stress on growth may be increased by heavy competition from shrub and herb layers.These results show the non-stationary nature of tree growth-climate associations and the linkage to for-est community structures.Vegetation components should be considered in future modeling and forecasting of forest dynamics in relation to climate changes.展开更多
The spatiotemporal variation in the altitudinal belts in the Kunlun Mountains reflects the natural geographical environment of the arid areas and is essential for constructing a dynamic change system of altitudinal be...The spatiotemporal variation in the altitudinal belts in the Kunlun Mountains reflects the natural geographical environment of the arid areas and is essential for constructing a dynamic change system of altitudinal belts in central and western China.In this study,we compiled the altitudinal belt spectra for three typical peaks(Amne Machin Mountain,Muztag Mountain,and Kongur Peak from east to west)using the Google Earth Engine(GEE),Landsat,and NASADEM.Considering historical vegetation surveys,we analyzed spatiotemporal patterns in the Kunlun Mountains for 2000,2010,and 2020.The results reveal that 1)the altitudinal belt spectra of the Kunlun Mountains exhibited a trend of complexity,simplicity,and complexity,with the number of altitudinal belts decreasing from six to three and then increasing to four from east to west.The dominant belt transitions from an alpine meadow belt to alpine desert steppe and montane desert steppe belts,excluding the nival belt.2)The upper limits of altitudinal belts are higher on sunny slopes than on shady slopes in the Kunlun Mountains,with an average altitudinal difference of 77.90 m.3)From 2000 to 2020,there was a widening trend in vegetation and desert belts,accompanied by glacier shrinkage.The range of glacier belts(nival and subnival)decreased by an average of 64 m,resulting in a 64 m rise in the upper limit of the alpine desert steppe belt in the Kunlun Mountains.This suggests drying and warming trends at high altitudes in the Kunlun Mountains in the study period.展开更多
Faeces were counted along horizontal, 5.0m wide strips at altitude intervals of 100m to examine the altitudinal ranging pattern of a band of black-and-white snub-nosed monkeys （Rhinopithecus bieti） at Nanren （99...Faeces were counted along horizontal, 5.0m wide strips at altitude intervals of 100m to examine the altitudinal ranging pattern of a band of black-and-white snub-nosed monkeys （Rhinopithecus bieti） at Nanren （99°04′E, 28°34′N, Baima Snow Mountain Nature Reverse, Yunnan, China） at four areas in each season between 2000-2001. Faeces were also counted along vertical, 2.5 m wide strips in one subvalley and on one subridge in each area. Monkeys used an altitudinal range of 3500-4300m, but preferred occupying the upper forest belt between 3900 and 4200m asl year-round, being at the highest altitude in summer, the lowest altitude in spring, and middle altitudes between autumn and winter （lower in winter than autumn）. Moreover, there were secondary peaks of lower altitude use （3700m） in spring and winter. In addition, more faeces were distributed in subvalleys than on subridges in winter, this suggests that monkeys tend to spend more time in subvalleys where there is less wind and fluctuating temperatures. The significant correlation between faecal density and lichen load might indicate that altitudinal distribution of food resources may act as an important factor affecting the monkeys＇ range. The monkeys might migrate to lower altitudes to eat young sprouts and leaves in spring, and to shelter from snowstorms in winter, possibly resulting in the secondary peaks at lower altitudes.展开更多
Fourteen plots were chosen along with the altitude of 1600-2600 m in Guandi Mountain, Shanxi Province, China to investigate all the trees in the plots, and the species diversity indexes were calculated for analyzing t...Fourteen plots were chosen along with the altitude of 1600-2600 m in Guandi Mountain, Shanxi Province, China to investigate all the trees in the plots, and the species diversity indexes were calculated for analyzing the effects of altitudinal gradient on plant species diversity. The results showed that the order of dominant species from low altitude to high altitude is as Quercus liaotungensis Koidz, Pinus tabulaeformis, Betula platyphylla, B. albo-sinensis, Picea wilsonii, P. Meyeri and Larix principis-rupprechtii. With altitude increasing, the average height and DBH of the arbor firstly increase and then decrease, the maximal height and DBH of the arbor present the unimodal variable trend, the maximal height of coniferous and broad-leaved trees firstly increases and then decreases, the maximal DBH of broad-leaf trees has no clear fluctuation, and the maximal DBH of the needle-leaf trees decreases gradually. Moreover, in middle altitudinal communities （the altitude of 1900-2200 m）, the maximal height and DBH of the needle-leaf trees are larger than these of broad-leaf trees. Both Shannon-wiener index and Margalef index of the communities present the unimodal variable trend, with a peak in the mid-altitude. Plant species diversity in the mid-altitude （1900-2200 m） is higher than that of the low altitude （1600-1900 m） and the high altitude （2200-2600 m）. In addition, the unimodal variable trend of α species diversity is clearly correlated with the altitudinal gradient. The change of plant species diversity is significantly correlated with the altitudinal gradient, and also related to the community structure, the community composition, the feature of species and the human disturbance.展开更多
In lowlands climate-specific processes due to weathering and erosion are dominant, whilst the geomorphology of mountains is dependent on the geologic-tectonic structure, i.e., the energy of erosion that increases acco...In lowlands climate-specific processes due to weathering and erosion are dominant, whilst the geomorphology of mountains is dependent on the geologic-tectonic structure, i.e., the energy of erosion that increases according to the vertical. The expression ＂extremely high mountains＂ has been established as the extreme of a continuous mountain classification. It has to be understood in terms of geomorphology, glaciology and vegetation. Correspondence of the planetary and hypsometric change of forms is of great value as synthetic explanation. It is confirmed with regard to vegetation, periglacial geomorphology and glaciology. Due to the world-wide reconstruction of the snowline its paleoclimatie importance increases, too. Apart from lower limits the periglacial and glacial altitudinal levels also show zones of optimum development and climatic upper limits in the highest mountains of the earth. According to the proportion of the altitudinal levels a classification as to arid, temperate and humid high mountains has been carried out.展开更多
Shrublands serve as an important component of terrestrial ecosystems, and play an important role in structure and functions of alpine ecosystem.Accurate estimation of biomass is critical to examination of the producti...Shrublands serve as an important component of terrestrial ecosystems, and play an important role in structure and functions of alpine ecosystem.Accurate estimation of biomass is critical to examination of the productivity of alpine ecosystems, due to shrubification under climate change in past decades.In this study, 14 experimental plots and 42 quadrates of the shrubs Potentilla fruticosa and Caragana jubata were selected along altitudes gradients from 3220 to 3650 m a.s.l.(above sea level) on semi-sunny and semi-shady slope in Hulu watershed of Qilian Mountains, China.The foliage, woody component and total aboveground biomass per quadrate were examined using a selective destructive method, then the biomass were estimated via allometric equations based on measured parameters for two shrub species.The results showed that C.jubata accounted for 1–3 times more biomass(480.98 g/m2) than P.fruticosa(191.21 g/m2).The aboveground biomass of both the shrubs varied significantly with altitudinal gradient(P<0.05).Woody component accounted for the larger proportion than foliage component in the total aboveground biomass.The biomass on semi-sunnyslopes(200.27 g/m2 and 509.07 g/m2) was greater than on semi-shady slopes(182.14 g/m2 and 452.89g/m2) at the same altitude band for P.fruticosa and C.jubata.In contrast, the foliage biomass on semi-shady slopes(30.50 g/m2) was greater than on semi-sunny slopes(27.51 g/m2) for two shrubs.Biomass deceased with increasing altitude for P.fruticosa, whereas C.jubata showed a hump-shaped pattern with altitude.Allometric equations were obtained from the easily descriptive parameters of height(H), basal diameter(D) and crown area(C) for biomass of C.jubata and P.fruticosa.Although the equations type and variables comprising of the best model varied among the species, all equations related to biomass were significant(P < 0.005), with determination coefficients(R2) ranging from 0.81 to 0.96.The allometric equations satisfied the requirements of the model, and can be used to estimate the regional scale biomass of P.fruticosa and C.jubata in alpine ecosystems of the Qilian Mountains.展开更多
The varied altitudinal gradient of climate and vegetation is further complicated by mass elevation effect(MEE), especially in high and extensive mountain regions. However, this effect and its implications for mountain...The varied altitudinal gradient of climate and vegetation is further complicated by mass elevation effect(MEE), especially in high and extensive mountain regions. However, this effect and its implications for mountain altitudinal belts have not been well studied until recently. This paper provides an overview of the research carried out in the past 5 years. MEE is virtually the heating effect of mountain massifs and can be defined as the temperature difference on a given elevation between inside and outside of a mountain mass. It can be digitally modelled with three factors of intra-mountain base elevation(MBE), latitude and hygrometric continentality; MBE usually acts as the primary factor for the magnitude of MEE and, to a great extent, could represent MEE. MEE leads to higher treelines in the interior than in the outside of mountain masses. It makes montane forests to grow at 4800–4900 m and snowlines to develop at about 6000 m in the southern Tibetan Plateau and the central Andes, and large areas of forests to live above 3500 m in a lot of high mountains of the world. The altitudinal distribution of global treelines can be modelled with high precision when taking into account MEE and the result shows that MEE contributes the most to treeline distribution pattern. Without MEE, forests could only develop upmost to about 3500 m above sea level and the world ecological pattern would be much simpler. The quantification of MEE should be further improved with higher resolution data and its global implications are to be further revealed.展开更多
This paper analyses the diversity and spatial pattern of the altitudinal belts in the Hengduan Mountains in China. A total of 7 types of base belts and 26 types of altitudinal belts are identified in the study region....This paper analyses the diversity and spatial pattern of the altitudinal belts in the Hengduan Mountains in China. A total of 7 types of base belts and 26 types of altitudinal belts are identified in the study region. The main altitudinal belt lines, such as forest line, the upper limit of dark coniferous forest and snow line, have similar latitudinal and longitudinal spatial patterns, namely, arched quadratic curve model with latitudes and concave quadratic curve model along longitudinal direction. These patterns can be together ealled as ＂Hyperbolic-paraboloid model＂, revealing the complexity and speciality of the environment and ecology in the study region. This result further validates the hypnosis of a common quadratic model for spatial pattern of mountain altitudinal belts proposed by the authors. The spatial pattern of altitudinal belts is closely related with moisture-related exposure effect in the Hengduan Mountains. Different combinations （spectra） of altitudinal belts and different base belt types appear in windward and leeward flanks and even in the same flanks of different ranges. This is closely related with the parallel mountain ranges of the Hengduan Mountains, which, at nearly right angle with the moving direction of prevailing moisture-laden air masses from west and east, hold up the warm and humid monsoon wind from moving into the core region and result in different moisture conditions in windward and leeward flanks. However, how to quantitatively describe the moisture-related exposure effect needs further study. In addition, the data quality and data accuracy at present also affect to some extent the result of quantitative modeling and should be improved with RS/GIS in the future.展开更多
The present study highlights the rich species diversity of higher plants in the Bhabha Valley of western Himalaya in India. The analysis of species diversity revealed that a total of 313 species of higher plants inhab...The present study highlights the rich species diversity of higher plants in the Bhabha Valley of western Himalaya in India. The analysis of species diversity revealed that a total of 313 species of higher plants inhabit the valley with a characteristic of moist alpine shrub vegetation. The herbaceous life forms dominate and increase with increasing altitude. The major representations are from the families Asteraceae, Rosaceae, Lamiaceae and Poaceae, suggesting thereby the alpine meadow nature of the study area. The effect of altitude on species diversity displays a hump-shaped curve which may be attributed to increase in habitat diversity at the median ranges and relatively less habitat diversity at higher altitudes. The anthropogenic pressure at lower altitudes results in low plant diversity towards the bottom of the valley with most of the species being exotic in nature. Though the plant diversity is less at higher altitudinal ranges, the uniqueness is relatively high with high species replacement rates. More than 90 % of variability in the species diversity could be explained using appropriate quantitative and statistical analysis along the altitudinal gradient. The valley harbours 18 threatened and 41 endemic species, most of which occur at higher altitudinal gradients due to habitat specificity.展开更多
Alpine timberline, as the ＂ecologica tion of scientists in many fields, especially in transition zone,＂ has long attracted the atten- recent years. Many unitary and dibasic fitting models have been developed to expl...Alpine timberline, as the ＂ecologica tion of scientists in many fields, especially in transition zone,＂ has long attracted the atten- recent years. Many unitary and dibasic fitting models have been developed to explore the relationship between timberline elevation and latitude or temperature. However, these models are usually on regional scale and could not be applied to other regions; on the other hand, hemispherical-scale and continental-scale models are usually based on about 100 timberline data and are necessarily low in precision. The present article collects 516 data sites of timberline, and takes latitude, continentality and mass elevation effect （MEE） as independent variables and timberline elevation as dependent variable to develop a ternary linear regression meteorological data released by WorldClim and model. Continentality is calculated using the mountain base elevation （as a proxy of mass elevation effect） is extracted on the basis of SRTM 90-meter resolution elevation data. The results show that the coefficient of determination （R2） of the linear model is as high as 0.904, and that the contribution rate of latitude, continentality and MEE to timberline elevation is 45.02% （p=0.000）, 6.04% （p=0.000） and 48.94% （p=0.000）, respectively. This means that MEE is simply the primary factor contributing to the elevation distribution of timberline on the continental and hemispherical scales. The contribution rate of MEE to timberline altitude dif- fers in different regions, e.g., 50.49% （p=0.000） in North America, 48.73% （p=0.000） in the eastern Eurasia, and 43.6% （p=0.000） in the western Eurasia, but it is usually very high.展开更多
Knowledge of vegetation distribution patterns is very important. Their relationships with topography and climate were explored through a geographically weighted regression （GWR） framework in a subtropical mountainou...Knowledge of vegetation distribution patterns is very important. Their relationships with topography and climate were explored through a geographically weighted regression （GWR） framework in a subtropical mountainous and hilly region, Minjiang River Basin of Fujian in China. The HJ-1 satellite image acquired on December 9, 2010 was utilized and NDVI index was calculated representing the range of vegetation greenness. Proper analysis units were achieved through segregation based on small sub-basins and altitudinal bands. Results indicated that the GWR model was more powerful than ordinary linear least square （OLS） regression in interpreting vegetation-environmental relationship, indicated by higher adjusted R2 and lower Akaike information criterion values. On one side, the OLS analysis revealed dominant positive influence from parameters of elevation and slope on vegetation distribution. On the other side, GWR analysis indicated that spatially, the parameters of topography had a very complex relationship with the vegetation distribution, as results of the various combinations of environmental factors, vegetation composition and also anthropogenic impact. The influences of elevation and slope generally decreased, from strongly positive to nearly zero, with increasing altitude and slope. Specially, most rapid changes of coefficients between NDVI and elevation or slope were observed in relatively flat and low-lying areas. This paper confirmed that the non-stationary analysis through the framework of GWR could lead to a better understanding of vegetation distribution in subtropical mountainous and hilly region. It was hoped that the proposed scale selection method combined with GWR framework would provide some guidelines on dealing with both spatial （horizontal） and altitudinal （vertical） non-stationarity in the dataset, and it could easily be applied in characterizing vegetation distribution patterns in other mountainous and hilly river basins and related research.展开更多
This study addressed the floral component traits and biomass allocation patterns of Gentiana hexaphylla as well as the relationships of these parameters along an elevation gradient(approximately 3700 m, 3800 m, 3900 m...This study addressed the floral component traits and biomass allocation patterns of Gentiana hexaphylla as well as the relationships of these parameters along an elevation gradient(approximately 3700 m, 3800 m, 3900 m, and 4000 m) on the eastern Qinghai-Tibet Plateau. The plant height, floral characteristics, and biomass allocation of G. hexaphylla were measured at different altitudes after field sampling, sorting, and drying. Plant height was significantly greater at 3700 m than that at other elevations. Flower length was significantly greater at 4000 m than that at other elevations, whereas the flower length at low elevations showed no significant differences. Corolla diameter increased with altitude, although the difference was not significant between 3800 m and 3900 m. Variations in biomass accumulation, including the aboveground, photosynthetic organ, flower and belowground biomasses, showed non-linear responses to changes in altitude. The aboveground and photosynthetic organ biomasses reached their lowest values at 4000 m, whereas the belowground and flower biomassreached minimum values at 3700 m. The sexual reproductive allocation of G. hexaphylla also increased with altitude, with a maximum observed at 4000 m. These results suggest that external environmental factors and altitudinal gradients as well as the biomass accumulation and allocation of G. hexaphylla play crucial roles in plant traits and significantly affect the ability of this species to adapt to harsh environments. The decreased number of flowers observed at higher altitudes may indicate a compensatory response for the lack of pollinators at high elevations, which is also suggested by the deformed flower shapes at high altitudes. In addition, the individual plant biomass(i.e., plant size) had significantly effect on flower length and corolla diameter. Based on the organ biomass results, the optimal altitude for G. hexaphylla in the eastern Qinghai-Tibet Plateau is 3800 m, where the plant exhibits minimum propagule biomass and asexual reproductive allocation.展开更多
Environmental variations and ontogeny may affect plant morphological traits and biomass allocation patterns that are related to the adjustments of plant ecological strategies. We selected 2-, 3-and 4-year-old Fritilla...Environmental variations and ontogeny may affect plant morphological traits and biomass allocation patterns that are related to the adjustments of plant ecological strategies. We selected 2-, 3-and 4-year-old Fritillaria unibracteata plants to explore the ontogenetic and altitudinal changes that impact their morphological traits(i.e., plant height, single leaf area,and specific leaf area) and biomass allocations [i.e.,biomass allocations of roots, bulbs, leaves, stems, and flowers] at relatively low altitudinal ranges(3400 m to 3600 m asl) and high altitudinal ranges(3600 m to4000 m asl). Our results indicated that plant height,root biomass allocation, and stem biomass allocation significantly increased during the process of individual growth and development, but single leaf area, specific leaf area, bulb biomass allocation, and leaf biomass allocation showed opposite trends.Furthermore, the impacts of altitudinal changes on morphological traits and biomass allocations had no significant differences at low altitude, except for single leaf area of 2-year-old plants. At high altitude,significantly reduced plant height, single leaf area and leaf biomass allocation for the 2-year-old plants,specific leaf area for the 2-and 4-year-old plants, and stem biomass allocation were found along altitudinal gradients. Significantly increased sexual reproductive allocation and relatively stable single leaf area and leaf biomass allocation were also observed for the 3-and 4-year-old plants. In addition, stable specific leaf area for the 3-year-old plants and root biomass allocation were recorded. These results suggested that the adaptive adjustments of alpine plants, in particular F. unibracteata were simultaneously influenced by altitudinal gradients and ontogeny.展开更多
In this paper, a digital identification method for the extraction of altitudinal belt spectra of montane natural belts is presented. Acquiring the sequential spectra of digital altitudinal belts in mountains at an acc...In this paper, a digital identification method for the extraction of altitudinal belt spectra of montane natural belts is presented. Acquiring the sequential spectra of digital altitudinal belts in mountains at an acceptable temporal frequency and over a large area requires extensive time and work if traditional methods of field investigation are to be used. Such being the case, often the altitudinal belts of a whole mountain or the belts at a regional scale are represented by single points. However, single points obviously cannot accurately reflect the spatial variety of altitudinal belts. In this context, a digital method was developed to extract the spectra of altitudinal belts from remote sensing data and SRTM DEM in the We.st Kunlun Mountains. By means of the 4km resolution SPOT-4 vegetation 10-day composite NDVI, the horizontal distribution of altitudinal belts were extracted through supervised classification, with a total classification accuracy of 72.23%. Then, a way of twice-scan was used to realize the automatic transition of horizontal maps to vertical belts. The classification results of remote-sensing data could thus be transformed automatically to sequential spectra of digital altitudinal belts. The upper and lower lines of the altitudinal belts were then extracted by vertical scanning of the belts. Relationships between the altitudinal belts based on the montane natural zones concerning vegetation types and the geomorphological altitudinal belts discussed. As a tentative method, were also the digital extraction method presented here is effective at digitally identifying altitudinal belts, and could be helpful in rapid information extraction over large-scale areas.展开更多
We analyzed the influence of climate change over the past 50 years on the radial growth of two tree species: Korean pine (Pinus koraiensis) and Yezo spruce (Picea jezoensis), located on Changbai Mountain, Northeast Ch...We analyzed the influence of climate change over the past 50 years on the radial growth of two tree species: Korean pine (Pinus koraiensis) and Yezo spruce (Picea jezoensis), located on Changbai Mountain, Northeast China, using a dendrochronology approach to understand factors that limit the altitude for tree species. Elevated temperatures increased the radial growth of Korean pine and decreased that of Yezo spruce. The positive response of tree growth to hydrothermal conditions was the key reason that the upper limit of elevation of Korean pine followed the temperature fluctuation pattern. Increased temperatures and precipitation and longer growing seasons accelerated Korean pine growth. As the temperature increased, correlations between Korean pine ring-width chronology and precipitation changed from negative to positive. In Yezo spruce, increasing monthly temperatures and inadequate precipitation during the middle and late parts of the growing season led to narrow growth rings, whereas decreasing monthly temperatures and sufficient precipitation during the late growing season promoted growth. Rising temperatures and adequate precipitation increases Korean pine growth, possibly elevating the upper range limit in altitude for this species. In contrast, Yezo spruce growth is negatively affected by warming temperatures and limited precipitation. Under future temperature increases and precipitation fluctuations, the upper limit altitude of Korean pine can reasonably be expected to shift upward and Yezo spruce downward.展开更多
The heating effect （or mass elevation effect, MEE） of the Tibetan Plateau （TP） is intense due to its massive body. Some studies have been undertaken on its role as the heat source in summer and its implications fo...The heating effect （or mass elevation effect, MEE） of the Tibetan Plateau （TP） is intense due to its massive body. Some studies have been undertaken on its role as the heat source in summer and its implications for Asian climate, but little has been known of the im- plications of its MEE for the distribution of mountain altitudinal belts （MABs）. Using air tem- perature data observed and remotely sensed data, MAB/treeline data, and ASTER GDEM data, this paper compares the height of MABs and alpine treelines in the main TP and the surrounding mountains/lowland and explains the difference from the point of view of MEE. The results demonstrate： 1） at same elevation, air temperature and the length of growing season gradually increase from the eastern edge to the interior TP, e.g., at 4500 m （corre- sponding to the mean altitude of the TP）, the monthly mean temperature is 3.58℃ higher （April） to 6.63℃ higher （June） in the interior plateau than in the Sichuan Basin; the 10℃ iso- therm for the warmest month goes upward from the edge to the interior of the plateau, at 4000 m in the Qilian Mts. and the eastern edges of the plateau, and up to 4600-5000 m in Lhasa and Zuogong; the warmth index at an altitude of 4500 m can be up to 15℃-month in the in- terior TP, but much lower at the eastern edges. 2） MABs and treeline follow a similar trend of rising inwards： dark-coniferous forest is 1000-1500 m higher and alpine steppe is about 700-900 m higher in the interior TP than at the eastern edges.展开更多
The compilation of 1:250,000 vegetation type map in the North-South transitional zone and 1:50,000 vegetation type maps in typical mountainous areas is one of the main tasks of Integrated Scientific Investigation of t...The compilation of 1:250,000 vegetation type map in the North-South transitional zone and 1:50,000 vegetation type maps in typical mountainous areas is one of the main tasks of Integrated Scientific Investigation of the North-South Transitional Zone of China.In the past,vegetation type maps were compiled by a large number of ground field surveys.Although the field survey method is accurate,it is not only time-consuming,but also only covers a small area due to the limitations of physical environment conditions.Remote sensing data can make up for the limitation of field survey because of its full coverage.However,there are still some difficulties and bottlenecks in the extraction of remote sensing information of vegetation types,especially in the automatic extraction.As an example of the compilation of 1:50,000 vegetation type map,this paper explores and studies the remote sensing extraction and mapping methods of vegetation type with medium and large scales based on mountain altitudinal belts of Taibai Mountain,using multi-temporal high resolution remote sensing data,ground survey data,previous vegetation type map and forest survey data.The results show that:1)mountain altitudinal belts can effectively support remote sensing classification and mapping of 1:50,000 vegetation type map in mountain areas.Terrain constraint factors with mountain altitudinal belt information can be generated by mountain altitudinal belts and 1:10,000 Digital Surface Model(DSM)data of Taibai Mountain.Combining the terrain constraint factors with multi-temporal and high-resolution remote sensing data,ground survey data and previous small-scale vegetation type map data,the vegetation types at all levels can be extracted effectively.2)The basic remote sensing interpretation and mapping process for typical mountains is interpretation of vegetation type-groups→interpretation of vegetation formation groups,formations and subformations→interpretation and classification of vegetation types&subtypes,which is a combination method of top-down method and bottom-up method,not the top-down or the bottom-up classification according to the level of mapping units.The results of this study provide a demonstration and scientific basis for the compilation of large and medium scale vegetation type maps.展开更多
Based on the framework of the geo-info spectra of montane altitudinal belts, this paper firstly reviews six classification systems for the spectra of mountain altitudinal belts in China and considers that detailed reg...Based on the framework of the geo-info spectra of montane altitudinal belts, this paper firstly reviews six classification systems for the spectra of mountain altitudinal belts in China and considers that detailed regional study of altitudinal belts is the key for reaching standardization and systemization of mountain altitudinal belts. Only can this further identify and resolve problems with the study of altitudinal belts. The factors forming the spectra of altitudinal belts are analyzed in the Tianshan Mountains of China, and a digital altitudinal belt system is constructed for the northern flank, southern flank, the heartland, and Ili valley in the west. The characteristics of each belt are revealed with a summarization of the pattern of areal differentiation of altitudinal belts.展开更多
Functional traits are predictors of plants in response to environmental stimuli. They represent specific functional adaptations to various environmental stresses. This study deals with the variation in plant functiona...Functional traits are predictors of plants in response to environmental stimuli. They represent specific functional adaptations to various environmental stresses. This study deals with the variation in plant functional traits along elevation gradient and land-use types in Sagarmatha National Park and Buffer Zone, Nepal. Two field investigations in April and September, 2011 were made to collect samples. Sampling was done from 2200 - 3800 m asl varying approx. 400 m. East and west facing aspects of each valley were chosen. In each aspect four land-use type categories including disturbed (cultivated land, exploited forest and meadow) and less disturbed natural forest were selected. A transect of 25 m long and 2.5 m wide was laid. Different eight traits of plants including lifeform, plant height, clonality, spinescence, leaf dry matter content, stem specific density, twig dry matter content and twig drying time were examined for 60 plant species belonging to 31 families, collected from 40 sampled plots. Nine different types of growth forms were recorded. Plant height of the investigated species ranged from 0.03 - 15 m. The stolon consisting species were dominant in exploited forests. Diversity of clonal species was more in meadow and non-clonal species were dominant in all the altitudes. Only eight species consisted of spines. In the disturbed land-use categories, we found high variation in a particular trait. Correlation analyses revealed the significant relationship (p < 0.01) among different traits. Herbs and shrubs were dominant at higher elevation and in disturbed land-use categories. Species from high altitude were mostly short basal herbs, while spinescence and tall trees were observed at lower altitudes. Species recorded in meadows and exploited forests showed high variation in traits due to disturbance mainly grazing, fire, litter collection and trampling. Altitudinal variation, climatic conditions and disturbance most strongly influence trait expression in the study area.展开更多
基金This research was funded by National Key Research and Development Program of China(No.2018YFA0605601)National Natural Science Foundation of China(No.42077417,41671042).
文摘A set of standard chronologies for tree-ring width(TRW),earlywood width(EWW)and latewood width(LWW)in Pinus tabuliformis Carr.along an altitudi-nal gradient(1450,1400,and 1350 m a.s.l.)on Baiyunshan Mountain,Central China to analyze the effect of varying temperature and precipitation on growth along the gradi-ent.Correlation analyses showed that at all three altitudes and the TRW and EWW chronologies generally had signifi-cant negative correlations with mean and maximum tem-peratures in the current April and May and with minimum temperatures in the prior July and August,but significant positive correlations with precipitation in the current May.Correlations were generally significantly negative between LWW chronologies and all temperatures in the prior July and August,indicating that the prior summer temperature had a strong lag effect on the growth of P.tabuliformis that increased with altitude.The correlation with the standard-ized precipitation evapotranspiration index(SPEI)confirmed that wet conditions in the current May promoted growth of TR and EW at all altitudes.Significant altitudinal differences were also found;at 1400 m,there were significant positive correlations between EWW chronologies and SPEI in the current April and significant negative correlations between LWW chronologies and SPEI in the current September,but these correlations were not significant at 1450 m.At 1350 m,there were also significant negative correlations between the TRW and the EWW chronologies and SPEI in the prior October and the current July and between LWW chronology and SPEI in the current August,but these cor-relations were not significant at 1400 m.Moving correlation results showed a stable response of EWW in relation to the SPEI in the current May at all three altitudes and of LWW to maximum temperature in the prior July-August at 1400 m from 2002 to 2018.The EWW chronology at 1400 m and the LWW chronology at 1450 m were identified as more suitable for climate reconstruction.These results provide a strong scientific basis for forest management decisions and climate reconstructions in Central China.
基金supported by the Second Tibetan Plateau Scientific Expedition and Research Program(2019QZKK0301)NationalNatural Science Foundation of China(32271886 and 42271074).
文摘Temporal changes in the relationship between tree growth and climate have been observed in numerous forests across the world.The patterns and the possible regu-lators(e.g.,forest community structure)of such changes are,however,not well understood.A vegetation survey and analyses of growth-climate relationships for Abies georgei var.Smithii(Smith fir)forests were carried along an altitudi-nal gradient from 3600 to 4200 m on Meili Snow Mountain,southeastern Tibetan Plateau.The results showed that the associations between growth and temperature have declined since the 1970s over the whole transect,while response to standardized precipitation-evapotranspiration indices(SPEI)strengthened in the mid-and lower-transect.Comparison between growth and vegetation data showed that tree growth was more sensitive to drought in stands with higher species richness and greater shrub cover.Drought stress on growth may be increased by heavy competition from shrub and herb layers.These results show the non-stationary nature of tree growth-climate associations and the linkage to for-est community structures.Vegetation components should be considered in future modeling and forecasting of forest dynamics in relation to climate changes.
基金Natural Science Foundation of Henan,No.232300420165,No.242300420218Integrated Scientific Investigation of the North-South Transitional Zone of China,No.2017FY100900。
文摘The spatiotemporal variation in the altitudinal belts in the Kunlun Mountains reflects the natural geographical environment of the arid areas and is essential for constructing a dynamic change system of altitudinal belts in central and western China.In this study,we compiled the altitudinal belt spectra for three typical peaks(Amne Machin Mountain,Muztag Mountain,and Kongur Peak from east to west)using the Google Earth Engine(GEE),Landsat,and NASADEM.Considering historical vegetation surveys,we analyzed spatiotemporal patterns in the Kunlun Mountains for 2000,2010,and 2020.The results reveal that 1)the altitudinal belt spectra of the Kunlun Mountains exhibited a trend of complexity,simplicity,and complexity,with the number of altitudinal belts decreasing from six to three and then increasing to four from east to west.The dominant belt transitions from an alpine meadow belt to alpine desert steppe and montane desert steppe belts,excluding the nival belt.2)The upper limits of altitudinal belts are higher on sunny slopes than on shady slopes in the Kunlun Mountains,with an average altitudinal difference of 77.90 m.3)From 2000 to 2020,there was a widening trend in vegetation and desert belts,accompanied by glacier shrinkage.The range of glacier belts(nival and subnival)decreased by an average of 64 m,resulting in a 64 m rise in the upper limit of the alpine desert steppe belt in the Kunlun Mountains.This suggests drying and warming trends at high altitudes in the Kunlun Mountains in the study period.
基金Doctoral Research Startup Grant at Southwest Forestry CollegeThe Chinese Academy of Science(KSCX2-1-03)
文摘Faeces were counted along horizontal, 5.0m wide strips at altitude intervals of 100m to examine the altitudinal ranging pattern of a band of black-and-white snub-nosed monkeys （Rhinopithecus bieti） at Nanren （99°04′E, 28°34′N, Baima Snow Mountain Nature Reverse, Yunnan, China） at four areas in each season between 2000-2001. Faeces were also counted along vertical, 2.5 m wide strips in one subvalley and on one subridge in each area. Monkeys used an altitudinal range of 3500-4300m, but preferred occupying the upper forest belt between 3900 and 4200m asl year-round, being at the highest altitude in summer, the lowest altitude in spring, and middle altitudes between autumn and winter （lower in winter than autumn）. Moreover, there were secondary peaks of lower altitude use （3700m） in spring and winter. In addition, more faeces were distributed in subvalleys than on subridges in winter, this suggests that monkeys tend to spend more time in subvalleys where there is less wind and fluctuating temperatures. The significant correlation between faecal density and lichen load might indicate that altitudinal distribution of food resources may act as an important factor affecting the monkeys＇ range. The monkeys might migrate to lower altitudes to eat young sprouts and leaves in spring, and to shelter from snowstorms in winter, possibly resulting in the secondary peaks at lower altitudes.
基金This paper was supported by Shanxi Natural Science Foun-dation (20001090)
文摘Fourteen plots were chosen along with the altitude of 1600-2600 m in Guandi Mountain, Shanxi Province, China to investigate all the trees in the plots, and the species diversity indexes were calculated for analyzing the effects of altitudinal gradient on plant species diversity. The results showed that the order of dominant species from low altitude to high altitude is as Quercus liaotungensis Koidz, Pinus tabulaeformis, Betula platyphylla, B. albo-sinensis, Picea wilsonii, P. Meyeri and Larix principis-rupprechtii. With altitude increasing, the average height and DBH of the arbor firstly increase and then decrease, the maximal height and DBH of the arbor present the unimodal variable trend, the maximal height of coniferous and broad-leaved trees firstly increases and then decreases, the maximal DBH of broad-leaf trees has no clear fluctuation, and the maximal DBH of the needle-leaf trees decreases gradually. Moreover, in middle altitudinal communities （the altitude of 1900-2200 m）, the maximal height and DBH of the needle-leaf trees are larger than these of broad-leaf trees. Both Shannon-wiener index and Margalef index of the communities present the unimodal variable trend, with a peak in the mid-altitude. Plant species diversity in the mid-altitude （1900-2200 m） is higher than that of the low altitude （1600-1900 m） and the high altitude （2200-2600 m）. In addition, the unimodal variable trend of α species diversity is clearly correlated with the altitudinal gradient. The change of plant species diversity is significantly correlated with the altitudinal gradient, and also related to the community structure, the community composition, the feature of species and the human disturbance.
文摘In lowlands climate-specific processes due to weathering and erosion are dominant, whilst the geomorphology of mountains is dependent on the geologic-tectonic structure, i.e., the energy of erosion that increases according to the vertical. The expression ＂extremely high mountains＂ has been established as the extreme of a continuous mountain classification. It has to be understood in terms of geomorphology, glaciology and vegetation. Correspondence of the planetary and hypsometric change of forms is of great value as synthetic explanation. It is confirmed with regard to vegetation, periglacial geomorphology and glaciology. Due to the world-wide reconstruction of the snowline its paleoclimatie importance increases, too. Apart from lower limits the periglacial and glacial altitudinal levels also show zones of optimum development and climatic upper limits in the highest mountains of the earth. According to the proportion of the altitudinal levels a classification as to arid, temperate and humid high mountains has been carried out.
基金funded by the National Natural Science Foundation of China(Grant Nos.91025011,91125013,41222001)the Project for Incubation of Specialists in Glaciology and Geocryology of National Natural Science Foundation of China(J1210003/J0109)
文摘Shrublands serve as an important component of terrestrial ecosystems, and play an important role in structure and functions of alpine ecosystem.Accurate estimation of biomass is critical to examination of the productivity of alpine ecosystems, due to shrubification under climate change in past decades.In this study, 14 experimental plots and 42 quadrates of the shrubs Potentilla fruticosa and Caragana jubata were selected along altitudes gradients from 3220 to 3650 m a.s.l.(above sea level) on semi-sunny and semi-shady slope in Hulu watershed of Qilian Mountains, China.The foliage, woody component and total aboveground biomass per quadrate were examined using a selective destructive method, then the biomass were estimated via allometric equations based on measured parameters for two shrub species.The results showed that C.jubata accounted for 1–3 times more biomass(480.98 g/m2) than P.fruticosa(191.21 g/m2).The aboveground biomass of both the shrubs varied significantly with altitudinal gradient(P<0.05).Woody component accounted for the larger proportion than foliage component in the total aboveground biomass.The biomass on semi-sunnyslopes(200.27 g/m2 and 509.07 g/m2) was greater than on semi-shady slopes(182.14 g/m2 and 452.89g/m2) at the same altitude band for P.fruticosa and C.jubata.In contrast, the foliage biomass on semi-shady slopes(30.50 g/m2) was greater than on semi-sunny slopes(27.51 g/m2) for two shrubs.Biomass deceased with increasing altitude for P.fruticosa, whereas C.jubata showed a hump-shaped pattern with altitude.Allometric equations were obtained from the easily descriptive parameters of height(H), basal diameter(D) and crown area(C) for biomass of C.jubata and P.fruticosa.Although the equations type and variables comprising of the best model varied among the species, all equations related to biomass were significant(P < 0.005), with determination coefficients(R2) ranging from 0.81 to 0.96.The allometric equations satisfied the requirements of the model, and can be used to estimate the regional scale biomass of P.fruticosa and C.jubata in alpine ecosystems of the Qilian Mountains.
基金National Natural Science Foundation of China,No.41421001,No.41571099,No.41030528
文摘The varied altitudinal gradient of climate and vegetation is further complicated by mass elevation effect(MEE), especially in high and extensive mountain regions. However, this effect and its implications for mountain altitudinal belts have not been well studied until recently. This paper provides an overview of the research carried out in the past 5 years. MEE is virtually the heating effect of mountain massifs and can be defined as the temperature difference on a given elevation between inside and outside of a mountain mass. It can be digitally modelled with three factors of intra-mountain base elevation(MBE), latitude and hygrometric continentality; MBE usually acts as the primary factor for the magnitude of MEE and, to a great extent, could represent MEE. MEE leads to higher treelines in the interior than in the outside of mountain masses. It makes montane forests to grow at 4800–4900 m and snowlines to develop at about 6000 m in the southern Tibetan Plateau and the central Andes, and large areas of forests to live above 3500 m in a lot of high mountains of the world. The altitudinal distribution of global treelines can be modelled with high precision when taking into account MEE and the result shows that MEE contributes the most to treeline distribution pattern. Without MEE, forests could only develop upmost to about 3500 m above sea level and the world ecological pattern would be much simpler. The quantification of MEE should be further improved with higher resolution data and its global implications are to be further revealed.
基金supported by the National Natural Science foundation of China (40971064)the Innovatioin Program of Chinese Academy of Sciences (No. kzcx2-yw-308)IGSNRR Youth Innovation Project,and the LREIS Self-determined Project
文摘This paper analyses the diversity and spatial pattern of the altitudinal belts in the Hengduan Mountains in China. A total of 7 types of base belts and 26 types of altitudinal belts are identified in the study region. The main altitudinal belt lines, such as forest line, the upper limit of dark coniferous forest and snow line, have similar latitudinal and longitudinal spatial patterns, namely, arched quadratic curve model with latitudes and concave quadratic curve model along longitudinal direction. These patterns can be together ealled as ＂Hyperbolic-paraboloid model＂, revealing the complexity and speciality of the environment and ecology in the study region. This result further validates the hypnosis of a common quadratic model for spatial pattern of mountain altitudinal belts proposed by the authors. The spatial pattern of altitudinal belts is closely related with moisture-related exposure effect in the Hengduan Mountains. Different combinations （spectra） of altitudinal belts and different base belt types appear in windward and leeward flanks and even in the same flanks of different ranges. This is closely related with the parallel mountain ranges of the Hengduan Mountains, which, at nearly right angle with the moving direction of prevailing moisture-laden air masses from west and east, hold up the warm and humid monsoon wind from moving into the core region and result in different moisture conditions in windward and leeward flanks. However, how to quantitatively describe the moisture-related exposure effect needs further study. In addition, the data quality and data accuracy at present also affect to some extent the result of quantitative modeling and should be improved with RS/GIS in the future.
文摘The present study highlights the rich species diversity of higher plants in the Bhabha Valley of western Himalaya in India. The analysis of species diversity revealed that a total of 313 species of higher plants inhabit the valley with a characteristic of moist alpine shrub vegetation. The herbaceous life forms dominate and increase with increasing altitude. The major representations are from the families Asteraceae, Rosaceae, Lamiaceae and Poaceae, suggesting thereby the alpine meadow nature of the study area. The effect of altitude on species diversity displays a hump-shaped curve which may be attributed to increase in habitat diversity at the median ranges and relatively less habitat diversity at higher altitudes. The anthropogenic pressure at lower altitudes results in low plant diversity towards the bottom of the valley with most of the species being exotic in nature. Though the plant diversity is less at higher altitudinal ranges, the uniqueness is relatively high with high species replacement rates. More than 90 % of variability in the species diversity could be explained using appropriate quantitative and statistical analysis along the altitudinal gradient. The valley harbours 18 threatened and 41 endemic species, most of which occur at higher altitudinal gradients due to habitat specificity.
基金National Natural Science Foundation of China,No.41030528No.40971064
文摘Alpine timberline, as the ＂ecologica tion of scientists in many fields, especially in transition zone,＂ has long attracted the atten- recent years. Many unitary and dibasic fitting models have been developed to explore the relationship between timberline elevation and latitude or temperature. However, these models are usually on regional scale and could not be applied to other regions; on the other hand, hemispherical-scale and continental-scale models are usually based on about 100 timberline data and are necessarily low in precision. The present article collects 516 data sites of timberline, and takes latitude, continentality and mass elevation effect （MEE） as independent variables and timberline elevation as dependent variable to develop a ternary linear regression meteorological data released by WorldClim and model. Continentality is calculated using the mountain base elevation （as a proxy of mass elevation effect） is extracted on the basis of SRTM 90-meter resolution elevation data. The results show that the coefficient of determination （R2） of the linear model is as high as 0.904, and that the contribution rate of latitude, continentality and MEE to timberline elevation is 45.02% （p=0.000）, 6.04% （p=0.000） and 48.94% （p=0.000）, respectively. This means that MEE is simply the primary factor contributing to the elevation distribution of timberline on the continental and hemispherical scales. The contribution rate of MEE to timberline altitude dif- fers in different regions, e.g., 50.49% （p=0.000） in North America, 48.73% （p=0.000） in the eastern Eurasia, and 43.6% （p=0.000） in the western Eurasia, but it is usually very high.
基金National Natural Science Foundation of China, No.41071207 No.41001254+2 种基金 Scientific Research Foundation for Returned Scholars, No.[20121940, Project from Ministry of Education of China and Science Foundation of Fujian Province, No.201210005 No.2012J01167 Research Foundations from Fuzhou University, No.2009-XQ-19
文摘Knowledge of vegetation distribution patterns is very important. Their relationships with topography and climate were explored through a geographically weighted regression （GWR） framework in a subtropical mountainous and hilly region, Minjiang River Basin of Fujian in China. The HJ-1 satellite image acquired on December 9, 2010 was utilized and NDVI index was calculated representing the range of vegetation greenness. Proper analysis units were achieved through segregation based on small sub-basins and altitudinal bands. Results indicated that the GWR model was more powerful than ordinary linear least square （OLS） regression in interpreting vegetation-environmental relationship, indicated by higher adjusted R2 and lower Akaike information criterion values. On one side, the OLS analysis revealed dominant positive influence from parameters of elevation and slope on vegetation distribution. On the other side, GWR analysis indicated that spatially, the parameters of topography had a very complex relationship with the vegetation distribution, as results of the various combinations of environmental factors, vegetation composition and also anthropogenic impact. The influences of elevation and slope generally decreased, from strongly positive to nearly zero, with increasing altitude and slope. Specially, most rapid changes of coefficients between NDVI and elevation or slope were observed in relatively flat and low-lying areas. This paper confirmed that the non-stationary analysis through the framework of GWR could lead to a better understanding of vegetation distribution in subtropical mountainous and hilly region. It was hoped that the proposed scale selection method combined with GWR framework would provide some guidelines on dealing with both spatial （horizontal） and altitudinal （vertical） non-stationarity in the dataset, and it could easily be applied in characterizing vegetation distribution patterns in other mountainous and hilly river basins and related research.
基金sponsored by the National Natural Science Foundation (Grant No.313705594, 31400389)China Postdoctoral ScienceFoundation under Grant (2014M552385)the International Science & Technology Cooperation Program of China (Grant No. 2013DFR90670)
文摘This study addressed the floral component traits and biomass allocation patterns of Gentiana hexaphylla as well as the relationships of these parameters along an elevation gradient(approximately 3700 m, 3800 m, 3900 m, and 4000 m) on the eastern Qinghai-Tibet Plateau. The plant height, floral characteristics, and biomass allocation of G. hexaphylla were measured at different altitudes after field sampling, sorting, and drying. Plant height was significantly greater at 3700 m than that at other elevations. Flower length was significantly greater at 4000 m than that at other elevations, whereas the flower length at low elevations showed no significant differences. Corolla diameter increased with altitude, although the difference was not significant between 3800 m and 3900 m. Variations in biomass accumulation, including the aboveground, photosynthetic organ, flower and belowground biomasses, showed non-linear responses to changes in altitude. The aboveground and photosynthetic organ biomasses reached their lowest values at 4000 m, whereas the belowground and flower biomassreached minimum values at 3700 m. The sexual reproductive allocation of G. hexaphylla also increased with altitude, with a maximum observed at 4000 m. These results suggest that external environmental factors and altitudinal gradients as well as the biomass accumulation and allocation of G. hexaphylla play crucial roles in plant traits and significantly affect the ability of this species to adapt to harsh environments. The decreased number of flowers observed at higher altitudes may indicate a compensatory response for the lack of pollinators at high elevations, which is also suggested by the deformed flower shapes at high altitudes. In addition, the individual plant biomass(i.e., plant size) had significantly effect on flower length and corolla diameter. Based on the organ biomass results, the optimal altitude for G. hexaphylla in the eastern Qinghai-Tibet Plateau is 3800 m, where the plant exhibits minimum propagule biomass and asexual reproductive allocation.
基金funded by the Natural Science Foundation Project of Sichuan Science and Technology Department (2018JY0305)Key Projects of the Natural Science Foundation of Sichuan Education Department (18ZA0002)
文摘Environmental variations and ontogeny may affect plant morphological traits and biomass allocation patterns that are related to the adjustments of plant ecological strategies. We selected 2-, 3-and 4-year-old Fritillaria unibracteata plants to explore the ontogenetic and altitudinal changes that impact their morphological traits(i.e., plant height, single leaf area,and specific leaf area) and biomass allocations [i.e.,biomass allocations of roots, bulbs, leaves, stems, and flowers] at relatively low altitudinal ranges(3400 m to 3600 m asl) and high altitudinal ranges(3600 m to4000 m asl). Our results indicated that plant height,root biomass allocation, and stem biomass allocation significantly increased during the process of individual growth and development, but single leaf area, specific leaf area, bulb biomass allocation, and leaf biomass allocation showed opposite trends.Furthermore, the impacts of altitudinal changes on morphological traits and biomass allocations had no significant differences at low altitude, except for single leaf area of 2-year-old plants. At high altitude,significantly reduced plant height, single leaf area and leaf biomass allocation for the 2-year-old plants,specific leaf area for the 2-and 4-year-old plants, and stem biomass allocation were found along altitudinal gradients. Significantly increased sexual reproductive allocation and relatively stable single leaf area and leaf biomass allocation were also observed for the 3-and 4-year-old plants. In addition, stable specific leaf area for the 3-year-old plants and root biomass allocation were recorded. These results suggested that the adaptive adjustments of alpine plants, in particular F. unibracteata were simultaneously influenced by altitudinal gradients and ontogeny.
基金funded by the National Natural Science Foundation of China (Grant No.40801045)the Knowledge Innovation Program of the Chinese Academy of Sciences (Grant No. kzcx2-yw-141)+2 种基金the Knowledge Innovation Program of the Chinese Academy of Sciences (Grant No. 0609211120)the National Natural Science Foundation of China (Grant No. 40801186)support from the postdoctoral project of UNAM
文摘In this paper, a digital identification method for the extraction of altitudinal belt spectra of montane natural belts is presented. Acquiring the sequential spectra of digital altitudinal belts in mountains at an acceptable temporal frequency and over a large area requires extensive time and work if traditional methods of field investigation are to be used. Such being the case, often the altitudinal belts of a whole mountain or the belts at a regional scale are represented by single points. However, single points obviously cannot accurately reflect the spatial variety of altitudinal belts. In this context, a digital method was developed to extract the spectra of altitudinal belts from remote sensing data and SRTM DEM in the We.st Kunlun Mountains. By means of the 4km resolution SPOT-4 vegetation 10-day composite NDVI, the horizontal distribution of altitudinal belts were extracted through supervised classification, with a total classification accuracy of 72.23%. Then, a way of twice-scan was used to realize the automatic transition of horizontal maps to vertical belts. The classification results of remote-sensing data could thus be transformed automatically to sequential spectra of digital altitudinal belts. The upper and lower lines of the altitudinal belts were then extracted by vertical scanning of the belts. Relationships between the altitudinal belts based on the montane natural zones concerning vegetation types and the geomorphological altitudinal belts discussed. As a tentative method, were also the digital extraction method presented here is effective at digitally identifying altitudinal belts, and could be helpful in rapid information extraction over large-scale areas.
基金financially supported by the Key Basic Research Project‘‘973’’(2010CB951301-5)the China Bureau of Foreign Experts and the Ministry of Education of China(111 Program,Grant 2008-B08044)
文摘We analyzed the influence of climate change over the past 50 years on the radial growth of two tree species: Korean pine (Pinus koraiensis) and Yezo spruce (Picea jezoensis), located on Changbai Mountain, Northeast China, using a dendrochronology approach to understand factors that limit the altitude for tree species. Elevated temperatures increased the radial growth of Korean pine and decreased that of Yezo spruce. The positive response of tree growth to hydrothermal conditions was the key reason that the upper limit of elevation of Korean pine followed the temperature fluctuation pattern. Increased temperatures and precipitation and longer growing seasons accelerated Korean pine growth. As the temperature increased, correlations between Korean pine ring-width chronology and precipitation changed from negative to positive. In Yezo spruce, increasing monthly temperatures and inadequate precipitation during the middle and late parts of the growing season led to narrow growth rings, whereas decreasing monthly temperatures and sufficient precipitation during the late growing season promoted growth. Rising temperatures and adequate precipitation increases Korean pine growth, possibly elevating the upper range limit in altitude for this species. In contrast, Yezo spruce growth is negatively affected by warming temperatures and limited precipitation. Under future temperature increases and precipitation fluctuations, the upper limit altitude of Korean pine can reasonably be expected to shift upward and Yezo spruce downward.
基金National Natural Science Foundation of China, No.41571099 No.41001278
文摘The heating effect （or mass elevation effect, MEE） of the Tibetan Plateau （TP） is intense due to its massive body. Some studies have been undertaken on its role as the heat source in summer and its implications for Asian climate, but little has been known of the im- plications of its MEE for the distribution of mountain altitudinal belts （MABs）. Using air tem- perature data observed and remotely sensed data, MAB/treeline data, and ASTER GDEM data, this paper compares the height of MABs and alpine treelines in the main TP and the surrounding mountains/lowland and explains the difference from the point of view of MEE. The results demonstrate： 1） at same elevation, air temperature and the length of growing season gradually increase from the eastern edge to the interior TP, e.g., at 4500 m （corre- sponding to the mean altitude of the TP）, the monthly mean temperature is 3.58℃ higher （April） to 6.63℃ higher （June） in the interior plateau than in the Sichuan Basin; the 10℃ iso- therm for the warmest month goes upward from the edge to the interior of the plateau, at 4000 m in the Qilian Mts. and the eastern edges of the plateau, and up to 4600-5000 m in Lhasa and Zuogong; the warmth index at an altitude of 4500 m can be up to 15℃-month in the in- terior TP, but much lower at the eastern edges. 2） MABs and treeline follow a similar trend of rising inwards： dark-coniferous forest is 1000-1500 m higher and alpine steppe is about 700-900 m higher in the interior TP than at the eastern edges.
基金National Natural Science Foundation of China,No.41871350,No.41571099Scientific and Technological Basic Resources Survey Project,No.2017FY 100900。
文摘The compilation of 1:250,000 vegetation type map in the North-South transitional zone and 1:50,000 vegetation type maps in typical mountainous areas is one of the main tasks of Integrated Scientific Investigation of the North-South Transitional Zone of China.In the past,vegetation type maps were compiled by a large number of ground field surveys.Although the field survey method is accurate,it is not only time-consuming,but also only covers a small area due to the limitations of physical environment conditions.Remote sensing data can make up for the limitation of field survey because of its full coverage.However,there are still some difficulties and bottlenecks in the extraction of remote sensing information of vegetation types,especially in the automatic extraction.As an example of the compilation of 1:50,000 vegetation type map,this paper explores and studies the remote sensing extraction and mapping methods of vegetation type with medium and large scales based on mountain altitudinal belts of Taibai Mountain,using multi-temporal high resolution remote sensing data,ground survey data,previous vegetation type map and forest survey data.The results show that:1)mountain altitudinal belts can effectively support remote sensing classification and mapping of 1:50,000 vegetation type map in mountain areas.Terrain constraint factors with mountain altitudinal belt information can be generated by mountain altitudinal belts and 1:10,000 Digital Surface Model(DSM)data of Taibai Mountain.Combining the terrain constraint factors with multi-temporal and high-resolution remote sensing data,ground survey data and previous small-scale vegetation type map data,the vegetation types at all levels can be extracted effectively.2)The basic remote sensing interpretation and mapping process for typical mountains is interpretation of vegetation type-groups→interpretation of vegetation formation groups,formations and subformations→interpretation and classification of vegetation types&subtypes,which is a combination method of top-down method and bottom-up method,not the top-down or the bottom-up classification according to the level of mapping units.The results of this study provide a demonstration and scientific basis for the compilation of large and medium scale vegetation type maps.
文摘Based on the framework of the geo-info spectra of montane altitudinal belts, this paper firstly reviews six classification systems for the spectra of mountain altitudinal belts in China and considers that detailed regional study of altitudinal belts is the key for reaching standardization and systemization of mountain altitudinal belts. Only can this further identify and resolve problems with the study of altitudinal belts. The factors forming the spectra of altitudinal belts are analyzed in the Tianshan Mountains of China, and a digital altitudinal belt system is constructed for the northern flank, southern flank, the heartland, and Ili valley in the west. The characteristics of each belt are revealed with a summarization of the pattern of areal differentiation of altitudinal belts.
文摘Functional traits are predictors of plants in response to environmental stimuli. They represent specific functional adaptations to various environmental stresses. This study deals with the variation in plant functional traits along elevation gradient and land-use types in Sagarmatha National Park and Buffer Zone, Nepal. Two field investigations in April and September, 2011 were made to collect samples. Sampling was done from 2200 - 3800 m asl varying approx. 400 m. East and west facing aspects of each valley were chosen. In each aspect four land-use type categories including disturbed (cultivated land, exploited forest and meadow) and less disturbed natural forest were selected. A transect of 25 m long and 2.5 m wide was laid. Different eight traits of plants including lifeform, plant height, clonality, spinescence, leaf dry matter content, stem specific density, twig dry matter content and twig drying time were examined for 60 plant species belonging to 31 families, collected from 40 sampled plots. Nine different types of growth forms were recorded. Plant height of the investigated species ranged from 0.03 - 15 m. The stolon consisting species were dominant in exploited forests. Diversity of clonal species was more in meadow and non-clonal species were dominant in all the altitudes. Only eight species consisted of spines. In the disturbed land-use categories, we found high variation in a particular trait. Correlation analyses revealed the significant relationship (p < 0.01) among different traits. Herbs and shrubs were dominant at higher elevation and in disturbed land-use categories. Species from high altitude were mostly short basal herbs, while spinescence and tall trees were observed at lower altitudes. Species recorded in meadows and exploited forests showed high variation in traits due to disturbance mainly grazing, fire, litter collection and trampling. Altitudinal variation, climatic conditions and disturbance most strongly influence trait expression in the study area.
